Basic Information

Faculty

• 農学部

Professional Memberships

• From Aug. 2024, To Present

  日本農芸化学会
• From Sep. 2023, To Present

  American Physiological Society
• From May 2017, To Present

  The Japan Neuroscience Society
• From Aug. 2014, To Jul. 2023

  The Japanese Biochemical Society
• From Aug. 2013, To Present

  The Molecular Biology Society of Japan
• From May 2012, To Present

  Japanese Society of Physical Fitness and Sports Medicine

Academic Degree

• 修士（人間・環境学）(京都大学)
• 博士（人間・環境学）(京都大学)

Research History

• From May 2021, To Mar. 2023

  Ritsumeikan University, Ritsumeikan-Global Innovation Research Organization, Visiting Researcher
• From Apr. 2021, To Present

  Kyoto University, Division of Food Science and Biotechnology, Graduate School of Agriculture, Assistant Professor
• From Apr. 2020, To Mar. 2021

  Ritsumeikan University, College of Gastronomy Management, Assistant
• From Apr. 2019, To Mar. 2020

  Ritsumeikan University, Research Organization of Science and Technology, Postdoctoral Fellow
• From Apr. 2016, To Mar. 2019

  Kyoto University, Laboratory of Sports and Exercise Medicine, Graduate School of Human and Environmental Studies, JSPS Research Fellow DC1

ID,URL

• ORCID ID

  0000-0002-3813-4958

• J-Global ID

  201901016868724683

• Researcher ID

  ABG-3699-2020

• Researcher number

  80844323

• External link

  http://www.sseiri.kais.kyoto-u.ac.jp/index.html

researchmap URL

• https://researchmap.jp/takumiyokokawa

Research

Research Interests

• 臓器連関
• exercise mimetics
• nutrition
• aging
• adipose tissue
• brain
• skeletal muscle
• exercise

Research Areas

• Life sciences, Neuroscience - general
• Life sciences, Metabolism and endocrinology
• Life sciences, Physiology
• Life sciences, Sports science
• Life sciences, Nutrition and health science

Papers

• 運動による白色脂肪組織の量的・機能的適応を制御する因子としての骨格筋AMPキナーゼの可能性

  横川 拓海; 江川 達郎; 林 達也

  デサントスポーツ科学, Feb. 2023, Peer-reviewed, Lead author

  
• CRTC1 in Mc4r-Expressing Cells Is Required for Peripheral Metabolism and Systemic Energy Homeostasis

  Haruka Miyamori; Takumi Yokokawa (co-first author); Motoki Miyakita; Kazuki Ozaki; Tsuyoshi Goto; Kazuo Inoue; Shigenobu Matsumura

  Diabetes, 12 Sep. 2024, Peer-reviewed, Lead author, Corresponding author

  
• Glycative stress inhibits hypertrophy and impairs cell membrane integrity in overloaded mouse skeletal muscle

  Tatsuro Egawa; Takeshi Ogawa; Takumi Yokokawa; Kohei Kido; Ryota Iyama; Haiyu Zhao; Eriko Kurogi; Katsumasa Goto; Tatsuya Hayashi

  Journal of Cachexia, Sarcopenia and Muscle, 04 Apr. 2024, Peer-reviewed

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• Altered expression of synaptic proteins and adhesion molecules in the hippocampus and cortex following the onset of diabetes in nonobese diabetic mice

  Takumi Yokokawa; Kohei Kido; Koji Sato; Tatsuya Hayashi; Satoshi Fujita

  Physiological Reports, 20 Apr. 2023, Peer-reviewed, Lead author, Corresponding author

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• Glycative stress and skeletal muscle dysfunctions: as an inducer of "Exercise-Resistance."

  Tatsuro Egawa; Takeshi Ogawa; Takumi Yokokawa; Kohei Kido; Mami Fujibayashi; Katsumasa Goto; Tatsuya Hayashi

  Glycative Stress Research, 31 Dec. 2022, Peer-reviewed

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• CRTC1 deficiency, specifically in melanocortin‐4 receptor‐expressing cells, induces hyperphagia, obesity, and insulin resistance

  Shigenobu Matsumura; Motoki Miyakita; Haruka Miyamori; Satomi Kyo; Fuka Ishikawa; Tsutomu Sasaki; Tomoki Jinno; Jin Tanaka; Kotomi Fujita; Takumi Yokokawa; Tsuyoshi Goto; Keiko Momma; Shigeo Takenaka; Kazuo Inoue

  The FASEB Journal, Dec. 2022, Peer-reviewed

  
• Association of serum brain-derived neurotrophic factor with hepatic enzymes, AST/ALT ratio, and FIB-4 index in middle-aged and older women

  Takumi Yokokawa; Shohei Sasaki; Kohei Sase; Naomi Yoshii; Jun Yasuda; Tatsuya Hayashi; Satoshi Fujita

  PLOS ONE, 23 Aug. 2022, Peer-reviewed, Lead author

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• The Effect of Glycation Stress on Skeletal Muscle

  Tatsuro Egawa; Kohei Kido; Takumi Yokokawa; Mami Fujibayashi; Katsumasa Goto; Tatsuya Hayashi

  Psychology and Patho-physiological Outcomes of Eating [Working Title], 05 May 2021

  
• TLR4-Mediated Inflammatory Responses Regulate Exercise-Induced Molecular Adaptations in Mouse Skeletal Muscle

  Haruna Fujiyoshi; Tatsuro Egawa; Eriko Kurogi; Takumi Yokokawa; Kohei Kido; Tatsuya Hayashi

  International Journal of Molecular Sciences, 07 Feb. 2022, Peer-reviewed

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• Relation between protein intake and resistance training-induced muscle hypertrophy in middle-aged women: a pilot study

  Jun Yasuda; Kao Murata; Tsubasa Hasegawa; Mana Yamamura; Sumiaki Maeo; Junya Takegaki; Nobuaki Tottori; Takumi Yokokawa; Risako Mori; Takuma Arimitsu; Shu Nishikori; Satoshi Fujita

  Nutrition, 25 Jan. 2022, Peer-reviewed

  
• Methylglyoxal reduces molecular responsiveness to 4 weeks of endurance exercise in mouse plantaris muscle

  Tatsuro Egawa; Takeshi Ogawa; Takumi Yokokawa; Kohei Kido; Katsumasa Goto; Tatsuya Hayashi

  Journal of Applied Physiology, 13 Jan. 2022, Peer-reviewed

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• Disruption of CRTC1 and CRTC2 in Sim1 cells strongly increases high-fat diet intake in female mice but has a modest impact on male mice

  Jin Tanaka; Fuka Ishikawa; Tomoki Jinno; Motoki Miyakita; Haruka Miyamori; Tsutomu Sasaki; Takumi Yokokawa; Tsuyoshi Goto; Kazuo Inoue; Shigenobu Matsumura

  PLOS ONE, 12 Jan. 2022, Peer-reviewed

  
• Caffeine increases myoglobin expression via the cyclic AMP pathway in L6 myotubes

  Takumi Yokokawa; Takeshi Hashimoto; Nobumasa Iwanaka

  Physiological Reports, May 2021, Peer-reviewed, Lead author, Corresponding author

  
• Involvement of receptor for advanced glycation end products in microgravity-induced skeletal muscle atrophy in mice

  Tatsuro Egawa; Kohei Kido; Takumi Yokokawa; Mami Fujibayashi; Katsumasa Goto; Tatsuya Hayashi

  Acta Astronautica, Nov. 2020, Peer-reviewed

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• Modified expression of vitamin D receptor and CYP27B1 in denervation-induced muscle atrophy

  Risako Mori; Takumi Yokokawa; Satoshi Fujita

  Biochemical and Biophysical Research Communications, Aug. 2020, Peer-reviewed

  
• Dehydroepiandrosterone activates 5′-adenosine monophosphate-activated protein kinase and suppresses lipid accumulation and adipocyte differentiation in 3T3-L1 cells

  Takumi Yokokawa; Koji Sato; Ryoko Narusawa; Kohei Kido; Risako Mori; Nobumasa Iwanaka; Tatsuya Hayashi; Takeshi Hashimoto

  Biochemical and Biophysical Research Communications, Jul. 2020, Peer-reviewed, Lead author, Corresponding author

  
• Muscle denervation reduces mitochondrial biogenesis and mitochondrial translation factor expression in mice

  Takumi Yokokawa; Risako Mori; Tadashi Suga; Tadao Isaka; Tatsuya Hayashi; Satoshi Fujita

  Biochemical and Biophysical Research Communications, Jun. 2020, Peer-reviewed, Lead author

  
• Enhanced skeletal muscle insulin sensitivity after acute resistance-type exercise is upregulated by rapamycin-sensitive mTOR complex 1 inhibition

  Kohei Kido; Kohei Sase; Takumi Yokokawa; Satoshi Fujita

  Scientific Reports, May 2020, Peer-reviewed

  
• The Protective Effect of Brazilian Propolis against Glycation Stress in Mouse Skeletal Muscle.

  Egawa T; Ohno Y; Yokoyama S; Yokokawa T; Tsuda S; Goto K; Hayashi T

  Foods (Basel, Switzerland), Sep. 2019, Peer-reviewed

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• A Lactate-based compound containing caffeine in addition to voluntary running exercise decreases subcutaneous fat mass and improves glucose metabolism in obese rats

  Hashimoto T; Yokokawa T; Narusawa R; Okada Y; Kawaguchi R; Higashida K

  Journal of Functional Foods, Mar. 2019, Peer-reviewed

  
• Exercise training increases CISD family protein expression in murine skeletal muscle and white adipose tissue.

  Yokokawa T; Kido K; Suga T; Sase K; Isaka T; Hayashi T; Fujita S

  Biochemical and biophysical research communications, Nov. 2018, Peer-reviewed, Lead author

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• Exercise-induced mitochondrial biogenesis coincides with the expression of mitochondrial translation factors in murine skeletal muscle.

  Yokokawa T; Kido K; Suga T; Isaka T; Hayashi T; Fujita S

  Physiological reports, Oct. 2018, Peer-reviewed, Lead author

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• Resistance training recovers attenuated APPL1 expression and improves insulin-induced akt signal activation in skeletal muscle of type 2 diabetic rats

  Kohei Kido; Satoru Ato; Takumi Yokokawa; Koji Sato; Satoshi Fujita

  American Journal of Physiology - Endocrinology and Metabolism, 01 Jun. 2018, Peer-reviewed

  
• Effect of resistance exercise under conditions of reduced blood insulin on AMPK alpha Ser485/491 inhibitory phosphorylation and AMPK pathway activation

  Kohei Kido; Takumi Yokokawa; Satoru Ato; Koji Sato; Satoshi Fujita

  AMERICAN JOURNAL OF PHYSIOLOGY-REGULATORY INTEGRATIVE AND COMPARATIVE PHYSIOLOGY, Aug. 2017, Peer-reviewed

  
• Acute bout of exercise induced prolonged muscle glucose transporter-4 translocation and delayed counter-regulatory hormone response in type 1 diabetes

  Koji Sato; Takeshi Nishijima; Takumi Yokokawa; Satoshi Fujita

  PLOS ONE, Jun. 2017, Peer-reviewed

  
• Acute resistance exercise-induced IGF1 expression and subsequent GLUT4 translocation

  Kohei Kido; Satoru Ato; Takumi Yokokawa; Yuhei Makanae; Koji Sato; Satoshi Fujita

  Physiological Reports, 01 Aug. 2016, Peer-reviewed

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• Dehydroepiandrosterone activates AMP kinase and regulates GLUT4 and PGC-1α expression in C2C12 myotubes

  Takumi Yokokawa; Koji Sato; Nobumasa Iwanaka; Hiroki Honda; Kazuhiko Higashida; Motoyuki Iemitsu; Tatsuya Hayashi; Takeshi Hashimoto

  Biochemical and Biophysical Research Communications, Jul. 2015, Peer-reviewed, Lead author

  
• Mixed lactate and caffeine compound increases satellite cell activity and anabolic signals for muscle hypertrophy.

  Yoshimi Oishi; Hayato Tsukamoto; Takumi Yokokawa; Keisuke Hirotsu; Mariko Shimazu; Kenji Uchida; Hironori Tomi; Kazuhiko Higashida; Nobumasa Iwanaka; Takeshi Hashimoto

  Journal of applied physiology (Bethesda, Md. : 1985), 15 Mar. 2015, Peer-reviewed

  
• Modest hypoxia significantly reduces triglyceride content and lipid droplet size in 3T3-L1 adipocytes

  Takeshi Hashimoto; Takumi Yokokawa; Yuriko Endo; Nobumasa Iwanaka; Kazuhiko Higashida; Sadayoshi Taguchi

  Biochemical and Biophysical Research Communications, Oct. 2013, Peer-reviewed

  
• Stimulation of G_s signaling in MC4R cells by DREADD increases energy expenditure, suppressing food intake, and increasing locomotor activity in mice

  Shigenobu Matsumura; Motoki Miyakita; Haruka Miyamori; Satomi Kyo; Daisuke Shima; Takumi Yokokawa; Fuka Ishikawa; Tsutomu Sasaki; Tomoki Jinno; Jin Tanaka; Tsuyoshi Goto; Keiko Momma; Kengo Ishihara; Rebecca Berdeaux; Kazuo Inoue

  American Journal of Physiology-Endocrinology and Metabolism, 28 Mar. 2022, Peer-reviewed

  

Presentations

• 生体適応の根幹を成す多臓器連関機構

  横川拓海

  第12回健康運動科学懇話会, 19 Oct. 2024, Invited

  
• 学習・情動制御におけるシナプス接着分子Arcadlinの役割

  岡﨑郁憲; 横川拓海; 澤田廣大; 井上和生

  2024年度日本農芸化学会関西支部大会（第532回講演会）―農芸化学会創立100周年・関西支部創立90周年記念―, 29 Sep. 2024

  
• 運動による脳機能調節に対する骨格筋AMPキナーゼの寄与

  澤田廣大; 横川拓海; 岡﨑郁憲; 江川達郎; 三浦進司; 井上和生; 林達也

  2024年度日本農芸化学会関西支部大会（第532回講演会）―農芸化学会創立100周年・関西支部創立90周年記念―, 29 Sep. 2024

  
• メラノコルチン経路による末梢組織代謝制御機構に関する新知見

  横川拓海

  第78回日本体力医学会大会, 02 Sep. 2024

  
• 成熟骨格筋におけるインスリン様成長因子1の欠損が骨格筋量および関連シグナル応答に及ぼす影響

  尾崎主基; 横川拓海; 江川達郎; 井山涼太; 林達也; 井上和生

  第38回日本体力医学会近畿地方会, 02 Mar. 2024

  
• Association between Serum 25-hydroxyvitamin D concentration and muscle mass and strength in Japanese elderly people

  Yamamura, M; Yasuda, J; Yokokawa, T; Fujita, S

  International Conference on Frailty and Sarcopenia Research 2020, 12 Mar. 2020

  
• 坐骨神経切除による骨格筋萎縮はビタミンD受容体およびCYP27B1のタンパク質発現量の増加を伴う

  森理紗子; 横川拓海; 藤田聡

  第42回日本分子生物学会年会, 06 Dec. 2019

  
• 骨格筋特異的IGF-I欠損が骨格筋ならびに全身性の代謝に及ぼす影響

  横川拓海; 木戸康平; 森理紗子; 佐瀬晃平; 増山律子; 林達也; 藤田聡

  第42回日本分子生物学会年会, 06 Dec. 2019

  
• マウスにおける経皮的電気刺激によるレジスタンス運動モデルの検討

  森理紗子; 横川拓海; 藤田聡

  第74回 日本体力医学会大会, Sep. 2019

  
• 脱神経がマウス骨格筋におけるミトコンドリア翻訳因子の発現量に及ぼす影響

  横川拓海; 森理紗子; 菅唯志; 伊坂忠夫; 林達也; 藤田聡

  第74回 日本体力医学会大会, Sep. 2019

  
• ラパマイシンは単回レジスタンス運動に伴う骨格筋のインスリン感受性亢進効果を高める

  木戸康平; 佐瀬晃平; 横川拓海; 藤田聡

  第74回 日本体力医学会大会, Sep. 2019

  
• Time course effects of voluntary running exercise on protein products of immediate early genes in murine hippocampus

  Yokokawa, T; Hayashi, T; Fujita, S

  24th Annual Congress of the European College of Sport Science, 04 Jul. 2019

  
• The influence of combination of Aurantiochytrium intake and resistance exercise on AMPK and mTORC1 signaling in murine skeletal muscle

  Mori, R; Yokokawa, T; Katamoto, Y; Ishihara, K; Iwata, O; Asayama, Y; Suzuki, K; Fujita, S

  24th Annual Congress of the European College of Sport Science, 03 Jul. 2019

  
• 自発性走運動がマウス海馬の最初期遺伝子およびシナプス分子に及ぼす影響の検討

  横川拓海; 林達也; 藤田聡

  第33回日本体力医学会近畿地方会, Feb. 2019

  
• オーランチオキトリウム抽出油の摂取と一過性レジスタンス運動の併用がマウス骨格筋のタンパク質合成シグナルに及ぼす影響

  森理紗子; 片本裕太; 横川拓海; 藤田聡

  第33回日本体力医学会近畿地方会, Feb. 2019

  
• Dysregulation of mitochondrial biogenesis and insulin signaling pathway in hippocampus of aged mice

  横川拓海; 木戸康平; 林達也; 藤田聡

  第41回日本分子生物学会年会, Nov. 2018

  
• Effect of vitamin D supplementation on resistance exercise-induced intramuscular vitamin D and muscle protein metabolism

  Katamoto, Y; Sase, K; Yokokawa, T; Fujita, S

  ACSM conference of Integrative Physiology of Exercise, 08 Sep. 2018

  
• 加齢マウスにおける不安行動の増大および海馬のシナプス分子の減少

  横川拓海; 守村直子; 高坂和芳; 木戸康平; 秦侑希; 山本かえで; 菅唯志; 田中秀和; 三品昌美; 林達也; 藤田聡

  2017年度生命科学系学会合同年次大会, Dec. 2017

  
• 電気けいれん療法はマウス海馬のmTORC1を活性化する

  横川拓海; 田中秀和; 井次陸; 高坂和芳; 林達也; 藤田聡

  第40回日本神経科学大会, Jul. 2017

  
• Chronic resistance training-induced increase in insulin-stimulated Akt Activation and GLUT4 translocation is associated with APPL1 expression in skeletal muscle of type 2 diabetic rat

  Kido, K; Ato, S; Nishimura, Y; Yokokawa, T; Sato, K; Fujita, S

  77th American Diabetes Association Scientific Sessions, Jun. 2017

  
• Reduced blood insulin stimulates resistance exercise-induced AMPK pathway activation concomitant with AMPKα Ser485/491 inhibitory phosphorylation in rat skeletal muscle

  Kido, K; Yokokawa, T; Ato, S; Sato, K; Fujita, S

  Cell Symposia Exercise Metabolism, 22 May 2017

  
• 慢性運動および除神経がマウス骨格筋におけるmitochondrial sirtuinsの発現量に及ぼす影響

  横川拓海; 菅唯志; 木戸康平; 阿藤聡; 林達也; 藤田聡; 伊坂忠夫

  第60回日本糖尿病学会年次学術集会, May 2017

  
• The Effect of Acute Resistance Exercise On Muscle Protein Synthesis in Atrophied Rat Skeletal Muscle After Unloading

  Ato, S; Kido, K; Miyake, T; Yokokawa, T; Fujita, S

  Experimental Biology 2017, 25 Apr. 2017

  
• Chronic exercise regulates the expression of mitochondrial sirtuins in murine skeletal muscle

  Yokokawa, T; Suga, T; Kido, K; Ato, S; Hayashi, T; Fujita, S; Isaka, T

  International Sports Science Network Forum in Nagano 2016, 10 Nov. 2016

  
• 自発的走運動はマウス海馬のmTORC1経路を活性化する

  横川拓海; 木戸康平; 阿藤聡; 田中秀和; 林達也; 藤田聡

  第39回 日本分子生物学会年会, Nov. 2016

  

External funds: Kakenhi

• Role of Arcadlin in exercise-induced enhancement of brain functions

  Grant-in-Aid for Early-Career Scientists

  Basic Section 59020:Sports sciences-related

  Kyoto University;Ritsumeikan University

  Takumi Yokokawa

  From 01 Apr. 2020, To 31 Mar. 2023, Project Closed

  Arcadlin;運動;海馬;情動;学習;脳;シナプス

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-20K19498/
• Role of skeletal muscle-derived IGF-I in exercise-induced enhancement of brain functions

  Grant-in-Aid for Research Activity Start-up

  0909:Sports sciences, physical education, health sciences, and related fields

  Kyoto University;Ritsumeikan University

  Takumi Yokokawa

  From 30 Aug. 2019, To 31 Mar. 2023, Project Closed

  インスリン様成長因子１;骨格筋;脳;臓器連関;情動;学習;インスリン成長因子1;海馬;インスリン様成長因子1;シナプス;インスリン様成長因子;運動;IGF-I

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19K24330/
• 運動に伴う骨格筋-脳の臓器連関を制御する分子としての骨格筋AMPキナーゼの可能性

  Grant-in-Aid for Challenging Research (Exploratory)

  Medium-sized Section 59:Sports sciences, physical education, health sciences, and related fields

  Kyoto University

  林 達也

  From 30 Jun. 2022, To 31 Mar. 2024, Granted

  骨格筋;脳;運動;AMPキナーゼ;マイオカイン;臓器連関

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-22K19750/
• 非酵素的翻訳後修飾の可視化から骨格筋の老化を「見る」

  Grant-in-Aid for Challenging Research (Pioneering)

  Medium-sized Section 59:Sports sciences, physical education, health sciences, and related fields

  Kyoto University

  江川 達郎

  From 30 Jun. 2022, To 31 Mar. 2025, Granted

  糖化;老化;イメージング;IMC;非酵素的翻訳後修飾;骨格筋

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-22K18413/
• 健康寿命延伸を目指したグロビン蛋白質生合成シグナルの解明

  Grant-in-Aid for Scientific Research (C)

  Basic Section 59040:Nutrition science and health science-related

  Kyoto Koka Women's University

  岩中 伸壮

  From 01 Apr. 2020, To 31 Mar. 2024, Granted

  グロビン蛋白質;シグナル伝達;運動;栄養;食品成分

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-20K11659/
• 運動による脳の健康増進における脂肪組織の寄与の解明

  Grant-in-Aid for Early-Career Scientists

  Basic Section 59020:Sports sciences-related

  Kyoto University

  横川 拓海

  From 01 Apr. 2023, To 31 Mar. 2026, Granted

  運動;脳;脂肪組織;臓器連関

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23K16695/
• 骨格筋糖化に着眼した個体老化の誘導機序解明

  Grant-in-Aid for Scientific Research (B)

  Basic Section 59020:Sports sciences-related

  Kyoto University

  林 達也

  From 01 Apr. 2023, To 31 Mar. 2026, Granted

  骨格筋;糖化ストレス;個体老化;臓器関連

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23H03283/
• 脳神経保護タンパク質に対する糖化ストレスの影響ならびに運動効果の検証

  Grant-in-Aid for Scientific Research (C)

  Basic Section 59020:Sports sciences-related

  Kyoto Koka Women's University

  岩中 伸壮

  From 01 Apr. 2024, To 31 Mar. 2027, Granted

  運動;糖化ストレス;ニューログロビン

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-24K14549/
• 骨格筋糖化に着眼した個体老化の誘導機序解明

  Grant-in-Aid for Scientific Research (B)

  Basic Section 59020:Sports sciences-related

  Kyoto University

  林 達也

  From 01 Apr. 2023, To 31 Mar. 2026, Granted

  骨格筋;糖化;老化;臓器連関;糖化ストレス;個体老化;臓器関連

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23K27973/

Education

Teaching subject(s)

• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Biochemistry and Microbiology

  7317, Fall, Faculty of Agriculture, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Advanced Course of Nuturitional Physiology II

  GA30, Fall, Graduate School of Agriculture, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar in Physiological Functions of Foods

  GF06, Spring, Graduate School of Agriculture, 1.5
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar in Physiological Functions of Foods

  GF06, Fall, Graduate School of Agriculture, 1.5
• From 01 Apr. 2024, To 31 Mar. 2025

  Experimental Course in Physiological Functions of Foods

  GG06, Spring, Graduate School of Agriculture, 3
• From 01 Apr. 2024, To 31 Mar. 2025

  Experimental Course in Physiological Functions of Foods

  GG06, Fall, Graduate School of Agriculture, 3
• From 01 Apr. 2024, To 31 Mar. 2025

  Laboratory Course in Molecular Biology

  3166, Spring, Faculty of Agriculture, 3
• From 01 Apr. 2023, To 31 Mar. 2024

  Practice of Basic Informatics (Faculty of Agriculture)

  T007, Fall, Institute for Liberal Arts and Sciences, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Biochemistry and Microbiology

  7317, Fall, Faculty of Agriculture, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Laboratory Course in Molecular Biology

  3166, Spring, Faculty of Agriculture, 3
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar in Physiological Functions of Foods2

  GB20, Year-long, Graduate School of Agriculture, 3
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar in Physiological Functions of Foods1

  GB19, Year-long, Graduate School of Agriculture, 3
• From 01 Apr. 2023, To 31 Mar. 2024

  Experimental Course in Physiological Functions of Foods1

  GC19, Year-long, Graduate School of Agriculture, 6
• From 01 Apr. 2023, To 31 Mar. 2024

  Experimental Course in Physiological Functions of Foods2

  GC20, Year-long, Graduate School of Agriculture, 6
• From 01 Apr. 2022, To 31 Mar. 2023

  Advanced Course of Nuturitional Physiology II

  GA30, Fall, Graduate School of Agriculture, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar in Physiological Functions of Foods1

  GB19, Year-long, Graduate School of Agriculture, 3
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Biochemistry and Microbiology

  7317, Fall, Faculty of Agriculture, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar in Physiological Functions of Foods2

  GB20, Year-long, Graduate School of Agriculture, 3
• From 01 Apr. 2022, To 31 Mar. 2023

  Experimental Course in Physiological Functions of Foods1

  GC19, Year-long, Graduate School of Agriculture, 6
• From 01 Apr. 2022, To 31 Mar. 2023

  Experimental Course in Physiological Functions of Foods2

  GC20, Year-long, Graduate School of Agriculture, 6
• From 01 Apr. 2022, To 31 Mar. 2023

  Laboratory Course in Molecular Biology

  3166, Spring, Faculty of Agriculture, 3
• From Apr. 2021, To Mar. 2022

  Laboratory Course in Molecular Biology

  Spring, 農学部
• From Apr. 2021, To Mar. 2022

  Seminar on Biochemistry and Microbiology

  Fall, 農学部
• From Apr. 2021, To Mar. 2022

  Experimental Course in Physiological Functions of Foods1

  Year-long, 農学研究科
• From Apr. 2021, To Mar. 2022

  Experimental Course in Physiological Functions of Foods2

  Year-long, 農学研究科
• From Apr. 2021, To Mar. 2022

  Seminar in Physiological Functions of Foods1

  Year-long, 農学研究科
• From Apr. 2021, To Mar. 2022

  Seminar in Physiological Functions of Foods2

  Year-long, 農学研究科