Basic Information

Affiliated programs (koza)

• Graduate School of Biostudies, 高次生命科学専攻　メカノセンシング生理学, 講師

Email Address

• Email Address

  goto.teppei.8ykyoto-u.ac.jp

Academic Degree

• 25 Mar. 2015

  名古屋大学博士（農学）
• 25 Mar. 2012

  名古屋大学修士（農学）

Academic Resume (Undergraduate School/Majors)

• 名古屋大学, 農学部資源生物科学科, 卒業

Research History

• From Apr. 2025, To Present

  Institute for Life and Medical Sciences, Kyoto University, Lab. of Mechanosensory Physiology, Lecturer
• From Oct. 2022, To Mar. 2025

  RIKEN, Center for Biosystems Dynamics Research, Postdoctoral researcher
• From Apr. 2019, To Sep. 2022

  RIKEN, Center for Biosystems Dynamics Research, Special Postdoctoral Researcher
• From Apr. 2016, To Mar. 2019

  National Institute for Physiological Sicences, Center for Genetic Analysis of Behavior, Postdoctoral research fellow
• From Apr. 2015, To Mar. 2016

  日本学術振興会, 特別研究員（DC2→PD）
• From Apr. 2014, To Mar. 2015

  日本学術振興会, 特別研究員（DC2）

ID,URL

• J-Global ID

  201801018784199953

researchmap URL

• https://researchmap.jp/teppeigoto

Research

Research Interests

• puberty onset
• kisspeptin
• interspecies chimera
• genome editing
• blastocyst complementation

Research Areas

• Life sciences, Zoological sciences
• Life sciences, Neuroscience - general
• Life sciences, Laboratory animal science
• Life sciences, Animals: biochemistry, physiology, behavioral science

Papers

• Dietary availability acutely influences puberty onset via a hypothalamic neural circuit

  Teppei Goto; Mitsue Hagihara; Satsuki Irie; Takaya Abe; Hiroshi Kiyonari; Kazunari Miyamichi

  Neuron, 24 Feb. 2025, Peer-reviewed, Lead author, Corresponding author

  
• Selective vulnerability of parvocellular oxytocin neurons in social dysfunction

  Masafumi Tsurutani; Teppei Goto; Mitsue Hagihara; Satsuki Irie; Kazunari Miyamichi

  Nature Communications, 06 Oct. 2024, Peer-reviewed

  
• Sex difference in developmental changes in visualized Kiss1 neurons in newly generated Kiss1-Cre rats.

  Koki Yamada; Mayuko Nagae; Tetsuya Mano; Hitomi Tsuchida; Safiullah Hazim; Teppei Goto; Makoto Sanbo; Masumi Hirabayashi; Naoko Inoue; Yoshihisa Uenoyama; Hiroko Tsukamura

  The Journal of reproduction and development, 28 Jul. 2023

  
• Dynamics of pulsatile activities of arcuate kisspeptin neurons in aging female mice

  Teppei Goto; Mitsue Hagihara; Kazunari Miyamichi

  eLife, 24 May 2023, Lead author, Corresponding author

  
• Characterization of homozygous Foxn1 mutations induced in rat embryos by different delivery forms of Cas9 nuclease.

  Teppei Goto; Kyoko Yogo; Shinichi Hochi; Masumi Hirabayashi

  Molecular biology reports, 28 Nov. 2022, Lead author

  
• Establishment of embryo transfer in the musk shrew (Suncus murinus).

  Naoko Inoue; Akinori Hotta; Teppei Goto; Masumi Hirabayashi; Yoshihisa Uenoyama; Hiroko Tsukamura

  The Journal of reproduction and development, 06 Oct. 2022

  
• 14-3-3 proteins stabilize LGI1-ADAM22 levels to regulate seizure thresholds in mice

  Norihiko Yokoi; Yuko Fukata; Kei Okatsu; Atsushi Yamagata; Yan Liu; Makoto Sanbo; Yuri Miyazaki; Teppei Goto; Manabu Abe; Hidetoshi Kassai; Kenji Sakimura; Dies Meijer; Masumi Hirabayashi; Shuya Fukai; Masaki Fukata

  Cell Reports, Dec. 2021

  Access this paper :

  
• Cellular and molecular mechanisms regulating the KNDy neuronal activities to generate and modulate GnRH pulse in mammals.

  Kana Ikegami; Youki Watanabe; Sho Nakamura; Teppei Goto; Naoko Inoue; Yoshihisa Uenoyama; Hiroko Tsukamura

  Frontiers in neuroendocrinology, 19 Nov. 2021

  
• Kisspeptin neurons as a key player bridging the endocrine system and sexual behavior in mammals.

  Sho Nakamura; Youki Watanabe; Teppei Goto; Kana Ikegami; Naoko Inoue; Yoshihisa Uenoyama; Hiroko Tsukamura

  Frontiers in neuroendocrinology, 29 Oct. 2021

  
• Blastocyst complementation using Prdm14-deficient rats enables efficient germline transmission and generation of functional mouse spermatids in rats.

  Toshihiro Kobayashi; Teppei Goto; Mami Oikawa; Makoto Sanbo; Fumika Yoshida; Reiko Terada; Naoko Niizeki; Naoyo Kajitani; Kanako Kazuki; Yasuhiro Kazuki; Shinichi Hochi; Hiromitsu Nakauchi; M Azim Surani; Masumi Hirabayashi

  Nature communications, 26 Feb. 2021, Lead author

  
• Direct evidence that KNDy neurons maintain gonadotropin pulses and folliculogenesis as the GnRH pulse generator

  Mayuko Nagae; Yoshihisa Uenoyama; Saki Okamoto; Hitomi Tsuchida; Kana Ikegami; Teppei Goto; Sutisa Majarune; Sho Nakamura; Makoto Sanbo; Masumi Hirabayashi; Kenta Kobayashi; Naoko Inoue; Hiroko Tsukamura

  Proceedings of the National Academy of Sciences, 02 Feb. 2021

  
• LGI1-ADAM22-MAGUK configures transsynaptic nanoalignment for synaptic transmission and epilepsy prevention.

  Yuko Fukata; Xiumin Chen; Satomi Chiken; Yoko Hirano; Atsushi Yamagata; Hiroki Inahashi; Makoto Sanbo; Hiromi Sano; Teppei Goto; Masumi Hirabayashi; Hans-Christian Kornau; Harald Prüss; Atsushi Nambu; Shuya Fukai; Roger A Nicoll; Masaki Fukata

  Proceedings of the National Academy of Sciences of the United States of America, 19 Jan. 2021

  Access this paper :

  
• Testosterone supplementation rescues spermatogenesis and in vitro fertilizing ability of sperm in Kiss1 knockout mice

  Teppei Goto; Masumi Hirabayashi; Youki Watanabe; Makoto Sanbo; Koichi Tomita; Naoko Inoue; Hiroko Tsukamura; Yoshihisa Uenoyama

  Endocrinology, 08 Jun. 2020, Peer-reviewed, Lead author

  
• Inducible Kiss1 knockdown in the hypothalamic arcuate nucleus suppressed pulsatile secretion of luteinizing hormone in male mice.

  Shiori Minabe; Sho Nakamura; Eri Fukushima; Marimo Sato; Kana Ikegami; Teppei Goto; Makoto Sanbo; Masumi Hirabayashi; Junko Tomikawa; Takuya Imamura; Naoko Inoue; Yoshihisa Uenoyama; Hiroko Tsukamura; Kei-Ichiro Maeda; Fuko Matsuda

  The Journal of reproduction and development, 26 Apr. 2020, Peer-reviewed

  
• Conditional kisspeptin neuron-specific Kiss1 knockout with newly generated Kiss1-floxed and Kiss1-Cre mice replicates a hypogonadal phenotype of global Kiss1 knockout mice.

  Kana Ikegami; Teppei Goto; Sho Nakamura; Youki Watanabe; Arisa Sugimoto; Sutisa Majarune; Kei Horihata; Mayuko Nagae; Junko Tomikawa; Takuya Imamura; Makoto Sanbo; Masumi Hirabayashi; Naoko Inoue; Kei-Ichiro Maeda; Hiroko Tsukamura; Yoshihisa Uenoyama

  The Journal of reproduction and development, 20 Apr. 2020, Peer-reviewed, Lead author

  
• Germline development in rat revealed by visualization and deletion of Prdm14

  Toshihiro Kobayashi; Hisato Kobayashi; Teppei Goto; Tomoya Takashima; Mami Oikawa; Hiroki Ikeda; Reiko Terada; Fumika Yoshida; Makoto Sanbo; Hiromitsu Nakauchi; Kazuki Kurimoto; Masumi Hirabayashi

  Development, 30 Jan. 2020, Peer-reviewed

  
• Pluripotent stem cell-derived organogenesis in the rat model system.

  Hirabayashi M; Goto T; Hochi S

  Transgenic research, Aug. 2019, Peer-reviewed

  
• Generation of pluripotent stem cell-derived mouse kidneys in Sall1-targeted anephric rats

  Goto T; Hara H; Sanbo M; Masaki H; Sato H; Yamaguchi T; Hochi S; Kobayashi T; Nakauchi H; Hirabayashi M

  Nature Communications, Feb. 2019, Peer-reviewed, Lead author

  
• Structural basis of epilepsy-related ligand–receptor complex LGI1–ADAM22

  Atsushi Yamagata; Yuri Miyazaki; Norihiko Yokoi; Hideki Shigematsu; Yusuke Sato; Sakurako Goto-Ito; Asami Maeda; Teppei Goto; Makoto Sanbo; Masumi Hirabayashi; Mikako Shirouzu; Yuko Fukata; Masaki Fukata; Shuya Fukai

  Nature Communications, Dec. 2018, Peer-reviewed

  
• An interspecies barrier to tetraploid complementation and chimera formation.

  Yamaguchi T; Sato H; Kobayashi T; Kato-Itoh M; Goto T; Hara H; Mizuno N; Yanagida A; Umino A; Hamanaka S; Suchy F; Masaki H; Ota Y; Hirabayashi M; Nakauchi H

  Scientific reports, Oct. 2018, Peer-reviewed

  
• Nylon Mesh Device for Vitrification of Large Quantities of Rat Pancreatic Islets

  Takahiro Yamanaka; Teppei Goto; Masumi Hirabayashi; Shinichi Hochi

  BIOPRESERVATION AND BIOBANKING, Oct. 2017, Peer-reviewed

  
• Evidence of involvement of neurone-glia/neurone-neurone communications via gap junctions in synchronised activity of KNDy neurones

  K. Ikegami; S. Minabe; N. Ieda; T. Goto; A. Sugimoto; S. Nakamura; N. Inoue; S. Oishi; A. D. Maturana; M. Sanbo; M. Hirabayashi; K-I Maeda; H. Tsukamura; Y. Uenoyama

  JOURNAL OF NEUROENDOCRINOLOGY, Jun. 2017, Peer-reviewed

  
• Interspecies organogenesis generates autologous functional islets

  Tomoyuki Yamaguchi; Hideyuki Sato; Megumi Kato-Itoh; Teppei Goto; Hiromasa Hara; Makoto Sanbo; Naoaki Mizuno; Toshihiro Kobayashi; Ayaka Yanagida; Ayumi Umino; Yasunori Ota; Sanae Hamanaka; Hideki Masaki; Sheikh Tamir Rashid; Masumi Hirabayashi; Hiromitsu Nakauchi

  NATURE, Feb. 2017, Peer-reviewed

  
• Haploid embryonic stem cell lines derived from androgenetic and parthenogenetic rat blastocysts

  Masumi Hirabayashi; Hiromasa Hara; Teppei Goto; Akiko Takizawa; Melinda R. Dwinell; Takahiro Yamanaka; Shinichi Hochi; Hiromitsu Nakauchi

  Journal of Reproduction and Development, 2017, Peer-reviewed

  
• Direct comparison of Cryotop (R) vitrification and Bicell (R) freezing on recovery of functional rat pancreatic islets

  Takahiro Yamanaka; Kazuya Tasbima; Rio Takahashi; Seiji Takashima; Teppei Goto; Masumi Hirabayashi; Shinichi Hochi

  CRYOBIOLOGY, Dec. 2016, Peer-reviewed

  
• Hypomorphic phenotype of Foxn1 gene-modified rats by CRISPR/Cas9 system

  Teppei Goto; Hiromasa Hara; Hiromitsu Nakauchi; Shinichi Hochi; Masumi Hirabayashi

  TRANSGENIC RESEARCH, Aug. 2016, Peer-reviewed, Lead author

  
• Rat Blastocysts from Nuclear Injection and Time-Lagged Enucleation and Their Commitment to Embryonic Stem Cells

  Hiromasa Hara; Teppei Goto; Akiko Takizawa; Makoto Sanbo; Howard J. Jacob; Toshihiro Kobayashi; Hiromitsu Nakauchi; Shinichi Hochi; Masumi Hirabayashi

  CELLULAR REPROGRAMMING, Apr. 2016, Peer-reviewed

  
• Molecular and Epigenetic Mechanism Regulating Hypothalamic Kiss1 Gene Expression in Mammals

  Yoshihisa Uenoyama; Junk Tomikawa; Naoko Inoue; Teppei Goto; Shiori Minabe; Nahoko Ieda; Sho Nakamura; Youki Watanabe; Kana Ikegami; Fuko Matsuda; Satoshi Ohkura; Kei-ichiro Maeda; Hiroko Tsukamura

  NEUROENDOCRINOLOGY, 2016, Peer-reviewed

  
• Knock-in of a Histone H2B-tdTomato Reporter Into the Rosa26 Locus Allows Visualization of Cell Nuclei in Rats

  Teppei Goto; Toshihiro Kobayashi; Hiromasa Hara; Makoto Sanbo; Shinichi Hochi; Hiromitsu Nakauchi; Masumi Hirabayashi

  MOLECULAR REPRODUCTION AND DEVELOPMENT, Dec. 2015, Peer-reviewed, Lead author

  
• Pharmacological and Morphological Evidence of AMPK-Mediated Energy Sensing in the Lower Brain Stem Ependymocytes to Control Reproduction in Female Rodents

  Shiori Minabe; Chikaya Deura; Kana Ikegami; Teppei Goto; Makoto Sanbo; Masumi Hirabayashi; Naoko Inoue; Yoshihisa Uenoyama; Kei-ichiro Maeda; Hiroko Tsukamura

  ENDOCRINOLOGY, Jun. 2015, Peer-reviewed

  
• Lack of Pulse and Surge Modes and Glutamatergic Stimulation of Luteinising Hormone Release in Kiss1 Knockout Rats

  Y. Uenoyama; S. Nakamura; Y. Hayakawa; K. Ikegami; Y. Watanabe; C. Deura; S. Minabe; J. Tomikawa; T. Goto; N. Ieda; N. Inoue; M. Sanbo; C. Tamura; M. Hirabayashi; K. -I. Maeda; H. Tsukamura

  JOURNAL OF NEUROENDOCRINOLOGY, Mar. 2015, Peer-reviewed

  
• Identification of Hypothalamic Arcuate Nucleus-Specific Enhancer Region of Kiss1 Gene in Mice

  Teppei Goto; Junko Tomikawa; Kana Ikegami; Shiori Minabe; Hitomi Abe; Tatsuya Fukanuma; Takuya Imamura; Kenji Takase; Makoto Sanbo; Koichi Tomita; Masumi Hirabayashi; Kei-ichiro Maeda; Hiroko Tsukamura; Yoshihisa Uenoyama

  MOLECULAR ENDOCRINOLOGY, Jan. 2015, Peer-reviewed, Lead author

  
• Derivation of Embryonic Stem Cell Lines from Parthenogenetically Developing Rat Blastocysts

  Masumi Hirabayashi; Teppei Goto; Chihiro Tamura; Makoto Sanbo; Hiromasa Hara; Megumi Kato-Itoh; Hideyuki Sato; Toshihiro Kobayashi; Hiromitsu Nakauchi; Shinichi Hochi

  STEM CELLS AND DEVELOPMENT, Jan. 2014, Peer-reviewed

  
• Effect of leukemia inhibitory factor and forskolin on establishment of rat embryonic stem cell lines

  Masumi Hirabayashi; Teppei Goto; Chihiro Tamura; Makoto Sanbo; Hiromasa Hara; Shinichi Hochi

  Journal of Reproduction and Development, 2014, Peer-reviewed

  
• Ability of tetraploid rat blastocysts to support fetal development after complementation with embryonic stem cells

  Masumi Hirabayashi; Chihiro Tamura; Makoto Sanbo; Teppei Goto; Megumi Kato-Itoh; Toshihiro Kobayashi; Hiromitsu Nakauchi; Shinichi Hochi

  MOLECULAR REPRODUCTION AND DEVELOPMENT, Jun. 2012, Peer-reviewed

  
• Epigenetic regulation of Kiss1 gene expression mediating estrogen-positive feedback action in the mouse brain

  Junko Tomikawa; Yoshihisa Uenoyama; Makiko Ozawa; Tatsuya Fukanuma; Kenji Takase; Teppei Goto; Hitomi Abe; Nahoko Ieda; Shiori Minabe; Chikaya Deura; Naoko Inoue; Makoto Sanbo; Koichi Tomita; Masumi Hirabayashi; Satoshi Tanaka; Takuya Imamura; Hiroaki Okamura; Kei-ichiro Maeda; Hiroko Tsukamura

  PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA, May 2012, Peer-reviewed

  

Misc.

• Histochemical and Cell Biological Studies of Transgenic Rats Prepared Using a Recombinase CreERT2-Introduced BAC Clone

  加藤幸雄; 上春浩貴; 後藤哲平; 吉田彩舟; 加藤たか子; 平林真澄

  明治大学農学部研究報告, Mar. 2018

  
• 膵臓欠損ラット体内で再生したマウスiPS細胞由来膵島の糖尿病誘発モデルマウスへの移植

  後藤 哲平; 山口 智之; 佐藤 秀征; 原 弘真; 小林 俊寛; 中内 啓光; 平林 真澄

  The Journal of Reproduction and Development, Sep. 2016

  
• 外径が異なるウシ卵核胞期卵丘卵子複合体ならびにラット膵ランゲルハンス島のガラス化耐性

  山中貴寛; 田島和弥; 高橋理央; 後藤哲平; 高島誠司; 高島誠司; 平林真澄; 保地眞一; 保地眞一

  Journal of Reproduction and Development, 2016

  
• コンディショナルKiss1KOマウスを用いたGnRHパルス発生機構の同定とKiss1可視化マウスin vitro系によるパルス発生分子メカニズムの解明

  池上 花奈; 美辺 詩織; 家田 菜穂子; 渡辺 雄貴; 後藤 哲平; 中村 翔; 井上 直子; 前多 敬一郎; 束村 博子; 上野山 賀久

  The Journal of Reproduction and Development, Sep. 2015

  

Presentations

• 膵臓欠損ラット体内で再生したマウスiPS細胞由来膵島の糖尿病誘発モデルマウスへの移植

  後藤 哲平; 山口 智之; 佐藤 秀征; 原 弘真; 小林 俊寛; 中内 啓光; 平林 真澄

  The Journal of Reproduction and Development, Sep. 2016, (一社)日本繁殖生物学会

  
• 膵臓欠損ラット体内で再生したマウスiPS細胞由来膵島の糖尿病誘発モデルマウスへの移植 (OR1-30)

  後藤哲平

  第109回日本繁殖生物学会, Sep. 2016

  
• Mouse thymus generated in Foxn1 mutant rats by blastocyst complementation

  Teppei Goto

  SSR 2016, Jul. 2016

  
• 外径が異なるウシ卵核胞期卵丘卵子複合体ならびにラット膵ランゲルハンス島のガラス化耐性

  山中貴寛; 田島和弥; 高橋理央; 後藤哲平; 高島誠司; 高島誠司; 平林真澄; 保地眞一; 保地眞一

  Journal of Reproduction and Development, 2016

  
• コンディショナルKiss1KOマウスを用いたGnRHパルス発生機構の同定とKiss1可視化マウスin vitro系によるパルス発生分子メカニズムの解明

  池上 花奈; 美辺 詩織; 家田 菜穂子; 渡辺 雄貴; 後藤 哲平; 中村 翔; 井上 直子; 前多 敬一郎; 束村 博子; 上野山 賀久

  The Journal of Reproduction and Development, Sep. 2015, (一社)日本繁殖生物学会

  
• CRISPR/Cas9法により作出したFoxN1遺伝子ノックアウトラットの解析 (P-91)

  後藤哲平

  第62回日本実験動物学会, May 2015

  
• Hypothalamic arcuate nuleus-specific enhancer for kisspeptin expression of female mice

  Teppei Goto

  WCRB2014, Sep. 2014

  
• Region-specific enhancer for Kiss1 expression in the hypothalamic arcuate nucleus of female mice (ZZ26)

  Teppei Goto

  Neuroscience 2013, Nov. 2013

  
• 遺伝子改変マウスを用いた Kiss1 発現におけるエンハンサーの同定 (30)

  後藤哲平

  第26回日本下垂体研究会学術集会, Aug. 2013

  
• Generation of Kiss1 Knockout Rats Evidencing the Role of Kisspeptin in Regulating Reproductive Function

  NAKAMURA Sho; UENOYAMA Yoshihisa; HAYAKAWA Yuki; IKEGAMI Kana; TOMIKAWA Junko; MINABE Shiori; GOTO Teppei; IEDA Nahoko; TAMURA Chihiro; SANBO Makoto; HIRABAYASHI Masumi; MAEDA Kei-ichiro; TSUKAMURA Hiroko

  The Journal of reproduction and development, 01 Feb. 2013

  
• Kiss1 Upstream Region Regulates Hypothalamic Arcuate Nucleus-Specific Kisspeptin Expression (P-34)

  Teppei Goto

  Kisspeptin2012, Nov. 2012

  
• 視床下部弓状核特異的なキスペプチン発現を制御する分子機構 (OR2-15)

  後藤哲平

  第105回日本繁殖生物学会, Sep. 2012

  
• 排卵制御中枢である前腹側室周囲核のキスペプチン発現制御機構 (O3-6-12)

  後藤哲平

  第85回日本内分泌学会, Apr. 2012

  
• Transgenic mice expressing GFP in two populations of kisspeptin in the hypothalamus

  Teppei Goto

  WCRB2011, Oct. 2011

  
• Kiss1遺伝子下流領域が前腹側室周囲核のキスペプチン発現に必須である (AW-1)

  後藤哲平

  第104回日本繁殖生物学会, Sep. 2011

  
• 緑色蛍光タンパク (GFP) によるキスペプチンニューロン可視化マウスの作出 (O3-8-15)

  後藤哲平

  第84回日本内分泌学会, Apr. 2011

  
• マウス胚の初期発生におけるキスペプチン-GPR54 系の役割

  後藤哲平

  第103回日本繁殖生物学会, Sep. 2010

  

Awards

• 09 Aug. 2013

  最優秀発表賞（日本下垂体研究会）
• 12 Mar. 2024

  理研桜舞賞（理化学研究所）
• 16 Sep. 2011

  口頭発表部門優秀発表賞（日本繁殖生物学会）

External funds: Kakenhi

• 性成熟期の生殖中枢に栄養不足を伝える神経ペプチドー受容体シグナリングの解明

  Grant-in-Aid for Scientific Research (C)

  Basic Section 46010:Neuroscience-general-related

  Institute of Physical and Chemical Research

  後藤 哲平

  From 01 Apr. 2024, To 31 Mar. 2027, Granted

  性成熟;キスペプチン;栄養不足

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-24K09651/
• Neural circuit basis for the onset of puberty by leptin signaling

  Grant-in-Aid for Early-Career Scientists

  Basic Section 46010:Neuroscience-general-related

  Institute of Physical and Chemical Research

  Teppei Goto

  From 01 Apr. 2019, To 31 Mar. 2021, Project Closed

  性成熟;キスペプチン;制限給餌;レプチン;逆行性トランスシナプス標識;トレーシング;神経回路

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19K16270/
• Changing activities of neurons controlling reproduction in menopausal transition and neural mechanism underlying hot flush

  Grant-in-Aid for Early-Career Scientists

  Basic Section 46010:Neuroscience-general-related

  Institute of Physical and Chemical Research

  Teppei Goto

  From 01 Apr. 2021, To 31 Mar. 2024, Project Closed

  更年期;老化;生殖中枢;イメージング;キスペプチン;ホットフラッシュ

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-21K15194/

Academic, Social Contribution

Social Contribution

• RIKEN BDR Summer School 2024

  Lecturer, Consultant

  RIKEN BDR, From 19 Aug. 2024, To 2024

  
• 生理研トレーニングコース

  Lecturer, Consultant

  生理学研究所, From 2012, To 2018