Basic Information

Email Address

• Email Address

  kometani.kohei.2mkyoto-u.ac.jp

Academic Degree

• 修士（生命科学）(京都大学)
• 博士（生命科学）(京都大学)

Academic Resume (Graduate Schools)

• 京都大学, 大学院生命科学研究科修士課程高次生命科学専攻, 修了
• 京都大学, 大学院生命科学研究科博士後期課程高次生命科学専攻, 修了

High School

• High School

  千葉県立　東葛飾高等学校

ID,URL

• J-Global ID

  200901033199796260

researchmap URL

• https://researchmap.jp/kohei_kometani

Research

Research Areas

• Life sciences, Immunology

Papers

• Altered immune function and hematologic malignancy in SPA-1-deficient mice

  D Ishida; K Kometani; A Tamura; S Li; M Hattori; N Minato

  IMMUNOLOGY 2004: AUTOIMMUNITY, GENETIC AND DEGENERATIVE DISORDERS, MALIGNANCIES, AND TRANSPLANTATION, 2004, Peer-reviewed

  
• CIN85 regulates the canonical NF-κB pathway and subsequent B cell responses.,

  Kometani K; Yamada T; Sasaki Y; Yokosuka T; Saito T; Rajewsky K; Ishiai M; Hikida M; Kurosaki T

  J. Exp. Med., 208(7):1447-57, 2011, 2011, Peer-reviewed

  
• Progressive differentiation toward the long-lived plasma cell compartment in the bone marrow

  Takuya Koike; Kentaro Fujii; Kohei Kometani; Noah S. Butler; Kenji Funakoshi; Shinya Yari; Junichi Kikuta; Masaru Ishii; Tomohiro Kurosaki; Wataru Ise

  Journal of Experimental Medicine, 06 Feb. 2023

  
• Identification of a T-bethi Quiescent Exhausted CD8 T Cell Subpopulation That Can Differentiate into TIM3+CX3CR1+ Effectors and Memory-like Cells.

  Saravanan Raju; Yu Xia; Bence Daniel; Kathryn E Yost; Elliot Bradshaw; Elena Tonc; Daniel J Verbaro; Kohei Kometani; Wayne M Yokoyama; Tomohiro Kurosaki; Ansuman T Satpathy; Takeshi Egawa

  Journal of immunology (Baltimore, Md. : 1950), 04 Jun. 2021

  
• Author Correction: BACH2 enforces the transcriptional and epigenetic programs of stem-like CD8+ T cells.

  Chen Yao; Guohua Lou; Hong-Wei Sun; Ziang Zhu; Yi Sun; Zeyu Chen; Daniel Chauss; E Ashley Moseman; Jun Cheng; Marc A D'Antonio; Wangke Shi; Junwei Shi; Kohei Kometani; Tomohiro Kurosaki; E John Wherry; Behdad Afzali; Luca Gattinoni; Yuwen Zhu; Dorian B McGavern; John J O'Shea; Pamela L Schwartzberg; Tuoqi Wu

  Nature immunology, Apr. 2021

  
• Selective expression of claudin-5 in thymic endothelial cells regulates the blood-thymus barrier and T-cell export.

  Takahiro Nagatake; Yan-Chun Zhao; Takeshi Ito; Masahiko Itoh; Kohei Kometani; Mikio Furuse; Azusa Saika; Eri Node; Jun Kunisawa; Nagahiro Minato; Yoko Hamazaki

  International immunology, 01 Mar. 2021

  
• BACH2 enforces the transcriptional and epigenetic programs of stem-like CD8+ T cells.

  Chen Yao; Guohua Lou; Hong-Wei Sun; Ziang Zhu; Yi Sun; Zeyu Chen; Daniel Chauss; E Ashley Moseman; Jun Cheng; Marc A D'Antonio; Wangke Shi; Junwei Shi; Kohei Kometani; Tomohiro Kurosaki; E John Wherry; Behdad Afzali; Luca Gattinoni; Yuwen Zhu; Dorian B McGavern; John J O'Shea; Pamela L Schwartzberg; Tuoqi Wu

  Nature immunology, Mar. 2021

  
• EBF1 and Pax5 safeguard leukemic transformation by limiting IL-7 signaling, Myc expression, and folate metabolism.

  Senthilkumar Ramamoorthy; Kohei Kometani; Josip S Herman; Marc Bayer; Sören Boller; Joy Edwards-Hicks; Haribaskar Ramachandran; Rui Li; Ramon Klein-Geltink; Erika L Pearce; Dominic Grün; Rudolf Grosschedl

  Genes & development, 01 Nov. 2020

  
• Bone Marrow Endothelial Cells Take Up Blood-Borne Immune Complexes via Fcγ Receptor IIb2 in an Erythropoietin-Dependent Manner.

  Takeshi Ito; Kohei Kometani; Nagahiro Minato; Yoko Hamazaki

  Journal of immunology (Baltimore, Md. : 1950), 15 Oct. 2020

  
• BACH2 drives quiescence and maintenance of resting Treg cells to promote homeostasis and cancer immunosuppression.

  Francis M Grant; Jie Yang; Rabab Nasrallah; James Clarke; Firas Sadiyah; Sarah K Whiteside; Charlotte J Imianowski; Paula Kuo; Panagiota Vardaka; Tihomir Todorov; Nordin Zandhuis; Ilinca Patrascan; David F Tough; Kohei Kometani; Robert Eil; Tomohiro Kurosaki; Klaus Okkenhaug; Rahul Roychoudhuri

  The Journal of experimental medicine, 07 Sep. 2020

  
• Attenuation of TCR-induced transcription by Bach2 controls regulatory T cell differentiation and homeostasis.

  Tom Sidwell; Yang Liao; Alexandra L Garnham; Ajithkumar Vasanthakumar; Renee Gloury; Jonas Blume; Peggy P Teh; David Chisanga; Christoph Thelemann; Fabian de Labastida Rivera; Christian R Engwerda; Lynn Corcoran; Kohei Kometani; Tomohiro Kurosaki; Gordon K Smyth; Wei Shi; Axel Kallies

  Nature communications, 14 Jan. 2020

  
• Inhibition of T cell activation and function by the adaptor protein CIN85.

  Mei Suen Kong; Akiko Hashimoto-Tane; Yusuke Kawashima; Machie Sakuma; Tadashi Yokosuka; Kohei Kometani; Reiko Onishi; Nick Carpino; Osamu Ohara; Tomohiro Kurosaki; Kia Kien Phua; Takashi Saito

  Science signaling, 05 Feb. 2019

  
• Revisiting the role of IRF3 in inflammation and immunity by conditional and specifically targeted gene ablation in mice.

  Hideyuki Yanai; Shiho Chiba; Sho Hangai; Kohei Kometani; Asuka Inoue; Yoshitaka Kimura; Takaya Abe; Hiroshi Kiyonari; Junko Nishio; Naoko Taguchi-Atarashi; Yu Mizushima; Hideo Negishi; Rudolf Grosschedl; Tadatsugu Taniguchi

  Proceedings of the National Academy of Sciences of the United States of America, 15 May 2018

  
• The adaptor molecule CD2AP in CD4 T cells modulates differentiation of follicular helper T cells during chronic LCMV infection.

  Saravanan Raju; Kohei Kometani; Tomohiro Kurosaki; Andrey S Shaw; Takeshi Egawa

  PLoS pathogens, May 2018

  
• KLRG1+ Effector CD8+ T Cells Lose KLRG1, Differentiate into All Memory T Cell Lineages, and Convey Enhanced Protective Immunity.

  Dietmar Herndler-Brandstetter; Harumichi Ishigame; Ryo Shinnakasu; Valerie Plajer; Carmen Stecher; Jun Zhao; Melanie Lietzenmayer; Lina Kroehling; Akiko Takumi; Kohei Kometani; Takeshi Inoue; Yuval Kluger; Susan M Kaech; Tomohiro Kurosaki; Takaharu Okada; Richard A Flavell

  Immunity, 17 Apr. 2018

  
• T Follicular Helper Cell-Germinal Center B Cell Interaction Strength Regulates Entry into Plasma Cell or Recycling Germinal Center Cell Fate.

  Wataru Ise; Kentaro Fujii; Katsuyuki Shiroguchi; Ayako Ito; Kohei Kometani; Kiyoshi Takeda; Eiryo Kawakami; Kazuo Yamashita; Kazuhiro Suzuki; Takaharu Okada; Tomohiro Kurosaki

  Immunity, 17 Apr. 2018

  
• The Role of BACH2 in T Cells in Experimental Malaria Caused by Plasmodium chabaudi chabaudi AS.

  Chelsea L Edwards; Marcela Montes de Oca; Fabian de Labastida Rivera; Rajiv Kumar; Susanna S Ng; Yulin Wang; Fiona H Amante; Kohei Kometani; Tomohiro Kurosaki; Tom Sidwell; Axel Kallies; Christian R Engwerda

  Frontiers in immunology, 2018

  
• Inflammatory responses induce an identity crisis of alveolar macrophages, leading to pulmonary alveolar proteinosis.

  Risa Ebina-Shibuya; Mitsuyo Matsumoto; Makoto Kuwahara; Kyoung-Jin Jang; Manabu Sugai; Yoshiaki Ito; Ryo Funayama; Keiko Nakayama; Yuki Sato; Naoto Ishii; Yasunobu Okamura; Kengo Kinoshita; Kohei Kometani; Tomohiro Kurosaki; Akihiko Muto; Masakazu Ichinose; Masakatsu Yamashita; Kazuhiko Igarashi

  The Journal of biological chemistry, 03 Nov. 2017

  
• Conversion of T cells to B cells by inactivation of polycomb-mediated epigenetic suppression of the B-lineage program.

  Tomokatsu Ikawa; Kyoko Masuda; Takaho A Endo; Mitsuhiro Endo; Kyoichi Isono; Yoko Koseki; Rinako Nakagawa; Kohei Kometani; Junichiro Takano; Yasutoshi Agata; Yoshimoto Katsura; Tomohiro Kurosaki; Miguel Vidal; Haruhiko Koseki; Hiroshi Kawamoto

  Genes & development, 15 Nov. 2016

  
• In vivo dendritic cell targeting cellular vaccine induces CD4+ Tfh cell-dependent antibody against influenza virus.

  Satoru Yamasaki; Kanako Shimizu; Kohei Kometani; Maki Sakurai; Masami Kawamura; Shin-Ichiro Fujii

  Scientific reports, 14 Oct. 2016

  
• Bach2-Batf interactions control Th2-type immune response by regulating the IL-4 amplification loop.

  Makoto Kuwahara; Wataru Ise; Mizuki Ochi; Junpei Suzuki; Kohei Kometani; Saho Maruyama; Maya Izumoto; Akira Matsumoto; Nobuaki Takemori; Ayako Takemori; Kenta Shinoda; Toshinori Nakayama; Osamu Ohara; Masaki Yasukawa; Tatsuya Sawasaki; Tomohiro Kurosaki; Masakatsu Yamashita

  Nature communications, 01 Sep. 2016

  
• Regulated selection of germinal-center cells into the memory B cell compartment.

  Ryo Shinnakasu; Takeshi Inoue; Kohei Kometani; Saya Moriyama; Yu Adachi; Manabu Nakayama; Yoshimasa Takahashi; Hidehiro Fukuyama; Takaharu Okada; Tomohiro Kurosaki

  Nature immunology, Jul. 2016

  
• Erratum: Responsiveness of B cells is regulated by the hinge region of IgD.

  Rudolf Übelhart; Eva Hug; Martina P Bach; Thomas Wossning; Marcus Dühren-von Minden; Anselm H C Horn; Dimitrios Tsiantoulas; Kohei Kometani; Tomohiro Kurosaki; Christoph J Binder; Heinrich Sticht; Lars Nitschke; Michael Reth; Hassan Jumaa

  Nature immunology, Jul. 2015

  
• Responsiveness of B cells is regulated by the hinge region of IgD.

  Rudolf Übelhart; Eva Hug; Martina P Bach; Thomas Wossning; Marcus Dühren-von Minden; Anselm H C Horn; Dimitrios Tsiantoulas; Kohei Kometani; Tomohiro Kurosaki; Christoph J Binder; Heinrich Sticht; Lars Nitschke; Michael Reth; Hassan Jumaa

  Nature immunology, May 2015

  
• Differentiation and maintenance of long-lived plasma cells.

  Kohei Kometani; Tomohiro Kurosaki

  Current opinion in immunology, Apr. 2015

  
• Memory B cells.

  Tomohiro Kurosaki; Kohei Kometani; Wataru Ise

  Nature reviews. Immunology, Mar. 2015

  
• The transcription repressors Bach2 and Bach1 promote B cell development by repressing the myeloid program.

  Ari Itoh-Nakadai; Reina Hikota; Akihiko Muto; Kohei Kometani; Miki Watanabe-Matsui; Yuki Sato; Masahiro Kobayashi; Atsushi Nakamura; Yuichi Miura; Yoko Yano; Satoshi Tashiro; Jiying Sun; Tomokatsu Ikawa; Kyoko Ochiai; Tomohiro Kurosaki; Kazuhiko Igarashi

  Nature immunology, Dec. 2014

  
• Memory B cells contribute to rapid Bcl6 expression by memory follicular helper T cells.

  Wataru Ise; Takeshi Inoue; James B McLachlan; Kohei Kometani; Masato Kubo; Takaharu Okada; Tomohiro Kurosaki

  Proceedings of the National Academy of Sciences of the United States of America, 12 Aug. 2014

  
• The Menin-Bach2 axis is critical for regulating CD4 T-cell senescence and cytokine homeostasis.

  Makoto Kuwahara; Junpei Suzuki; Soichi Tofukuji; Takeshi Yamada; Makoto Kanoh; Akira Matsumoto; Saho Maruyama; Kohei Kometani; Tomohiro Kurosaki; Osamu Ohara; Toshinori Nakayama; Masakatsu Yamashita

  Nature communications, 02 Apr. 2014

  
• Defective immune responses in mice lacking LUBAC-mediated linear ubiquitination in B cells.

  Yoshiteru Sasaki; Soichi Sano; Masaki Nakahara; Shigeo Murata; Kohei Kometani; Yuichi Aiba; Shinji Sakamoto; Yoshihiro Watanabe; Keiji Tanaka; Tomohiro Kurosaki; Kazuhiro Iwai

  The EMBO journal, 11 Sep. 2013

  
• Repression of the transcription factor Bach2 contributes to predisposition of IgG1 memory B cells toward plasma cell differentiation.

  Kohei Kometani; Rinako Nakagawa; Ryo Shinnakasu; Tomohiro Kaji; Andrei Rybouchkin; Saya Moriyama; Koji Furukawa; Haruhiko Koseki; Toshitada Takemori; Tomohiro Kurosaki

  Immunity, 25 Jul. 2013

  
• Bach2 maintains T cells in a naive state by suppressing effector memory-related genes.

  Shin-ichi Tsukumo; Midori Unno; Akihiko Muto; Arata Takeuchi; Kohei Kometani; Tomohiro Kurosaki; Kazuhiko Igarashi; Takashi Saito

  Proceedings of the National Academy of Sciences of the United States of America, 25 Jun. 2013

  
• Gene expression profiling of neural stem cells and identification of regulators of neural differentiation during cortical development.

  Toshiyuki Ohtsuka; Hiromi Shimojo; Mitsuhiro Matsunaga; Naoki Watanabe; Kohei Kometani; Nagahiro Minato; Ryoichiro Kageyama

  Stem cells (Dayton, Ohio), Nov. 2011

  
• CIN85 drives B cell responses by linking BCR signals to the canonical NF-kappaB pathway.

  Kohei Kometani; Takayuki Yamada; Yoshiteru Sasaki; Tadashi Yokosuka; Takashi Saito; Klaus Rajewsky; Masamichi Ishiai; Masaki Hikida; Tomohiro Kurosaki

  The Journal of experimental medicine, 04 Jul. 2011

  
• Tangential migration and proliferation of intermediate progenitors of GABAergic neurons in the mouse telencephalon.

  Shengxi Wu; Shigeyuki Esumi; Keisuke Watanabe; Jing Chen; Kouichi C Nakamura; Kazuhiro Nakamura; Kouhei Kometani; Nagahiro Minato; Yuchio Yanagawa; Kaori Akashi; Kenji Sakimura; Takeshi Kaneko; Nobuaki Tamamaki

  Development (Cambridge, England), Jun. 2011

  
• ERKs induce expression of the transcriptional repressor Blimp-1 and subsequent plasma cell differentiation.

  Tomoharu Yasuda; Kohei Kometani; Noriko Takahashi; Yuuki Imai; Yuichi Aiba; Tomohiro Kurosaki

  Science signaling, 19 Apr. 2011

  
• Unique properties of memory B cells of different isotypes.

  Tomohiro Kurosaki; Yuichi Aiba; Kohei Kometani; Saya Moriyama; Yoshimasa Takahashi

  Immunological reviews, Sep. 2010

  
• Preferential localization of IgG memory B cells adjacent to contracted germinal centers.

  Yuichi Aiba; Kohei Kometani; Megumi Hamadate; Saya Moriyama; Asako Sakaue-Sawano; Michio Tomura; Hervé Luche; Hans Jörg Fehling; Rafael Casellas; Osami Kanagawa; Atsushi Miyawaki; Tomohiro Kurosaki

  Proceedings of the National Academy of Sciences of the United States of America, 06 Jul. 2010

  
• Essential role of Rap signal in pre-TCR-mediated beta-selection checkpoint in alphabeta T-cell development.

  Kohei Kometani; Masaki Moriyama; Yasuhiro Nakashima; Yoshinori Katayama; Shu-Fang Wang; Sho Yamasaki; Takashi Saito; Masakazu Hattori; Nagahiro Minato

  Blood, 01 Dec. 2008

  
• Bone marrow pre-B expansion by SL/Kh-Bomb1 locus: not sufficient for lymphomagenesis.

  Takuya Hiratsuka; Tatsuaki Tsuruyama; Richard Kaszynski; Kohei Kometani; Nagahiro Minato; Takuro Nakamura; Keiji Tamaki; Hiroshi Hiai

  Leukemia research, Feb. 2008

  
• SMAD4-deficient intestinal tumors recruit CCR1+ myeloid cells that promote invasion.

  Takanori Kitamura; Kohei Kometani; Hiroki Hashida; Akihiro Matsunaga; Hiroyuki Miyoshi; Hisahiro Hosogi; Masahiro Aoki; Masanobu Oshima; Masakazu Hattori; Arimichi Takabayashi; Nagahiro Minato; Makoto M Taketo

  Nature genetics, Apr. 2007

  
• Regulation of immune responses and hematopoiesis by the Rap1 signal.

  Nagahiro Minato; Kohei Kometani; Masakazu Hattori

  Advances in immunology, 2007

  
• Role of SPA-1 in phenotypes of chronic myelogenous leukemia induced by BCR-ABL-expressing hematopoietic progenitors in a mouse model.

  Kohei Kometani; Misayo Aoki; Shin Kawamata; Yoriko Shinozuka; Takumi Era; Masafumi Taniwaki; Masakazu Hattori; Nagahiro Minato

  Cancer research, 15 Oct. 2006

  
• Pyramidal neurons of upper cortical layers generated by NEX-positive progenitor cells in the subventricular zone.

  Sheng-Xi Wu; Sandra Goebbels; Kouichi Nakamura; Kazuhiro Nakamura; Kouhei Kometani; Nagahiro Minato; Takeshi Kaneko; Klaus-Armin Nave; Nobuaki Tamamaki

  Proceedings of the National Academy of Sciences of the United States of America, 22 Nov. 2005

  
• Rap1 and SPA-1 in hematologic malignancy.

  Kohei Kometani; Daisuke Ishida; Masakazu Hattori; Nagahiro Minato

  Trends in molecular medicine, Aug. 2004, Peer-reviewed, Lead author

  
• Myeloproliferative stem cell disorders by deregulated Rap1 activation in SPA-1-deficient mice.

  Daisuke Ishida; Kohei Kometani; Hailin Yang; Kiyokazu Kakugawa; Kyoko Masuda; Kazuhiro Iwai; Misao Suzuki; Shigeyoshi Itohara; Tatsutoshi Nakahata; Hiroshi Hiai; Hiroshi Kawamoto; Masakazu Hattori; Nagahiro Minato

  Cancer cell, Jul. 2003

  

Misc.

• Gene expression profiling of sorted neural stem cells during cortical development

  T. Ohtsuka; H. Shimojo; K. Kometani; N. Minato; R. Kageyama

  MECHANISMS OF DEVELOPMENT, Sep. 2005, Peer-reviewed

  
• エフェクターCD8陽性T細胞の分化の可塑性によりメモリーCD8陽性T細胞の多様性が形成される

  石亀晴道; 石亀晴道; HERNDLER-BRANDS Dietmer; 新中須亮; 井上毅; 米谷耕平; 黒崎知博; 黒崎知博; FLAVELL Richard; 岡田峰陽

  日本インターフェロン・サイトカイン学会学術集会抄録集, 2018

  
• B細胞内のシグナル伝達機構(第2回) B細胞シグナル伝達におけるアダプター分子の役割

  米谷 耕平

  炎症と免疫, Jun. 2015

  
• メモリーB細胞から形質細胞への分化と転写因子

  米谷 耕平

  臨床免疫・アレルギー科, Dec. 2014

  
• 転写因子Bach2によるリンパ球制御

  米谷 耕平; 黒崎 知博

  感染・炎症・免疫, Apr. 2014

  
• 【免疫学のブラックボックス 免疫記憶の制御と疾患治療 病原体を記憶する細胞と、アレルギー・癌に対するワクチンの有効性】(第2章)B細胞免疫記憶の総合的理解 記憶B細胞からプラズマ細胞への分化制御

  米谷 耕平; 黒崎 知博

  実験医学, Nov. 2011

  
• T細胞の初期分化 胸腺内T細胞分化β-selectionにおけるRap1シグナルの関与

  森山 正樹; 米谷 耕平; 山崎 晶; 斉藤 隆; 服部 雅一; 湊 長博

  日本免疫学会総会・学術集会記録, Nov. 2006

  
• 胸腺内T細胞分化における低分子量G蛋白質,Rap1の役割

  米谷 耕平; 新田 剛; 服部 雅一; 竹田 潤二; 高浜 洋介; 湊 長博

  日本免疫学会総会・学術集会記録, Nov. 2004

  
• SPA-1遺伝子破壊マウスにおける骨髄増殖性疾患の発症

  石田 大輔; 米谷 耕平; 服部 雅一; 湊 長博

  日本癌学会総会記事, Sep. 2004

  
• 慢性骨髄性白血病発症におけるSPA-1の関与

  米谷 耕平; 服部 雅一; 湊 長博

  日本癌学会総会記事, Sep. 2004

  
• Spa-1分子によるB1細胞機能制御

  石田 大輔; 米谷 耕平; 浜崎 洋子; 服部 雅一; 湊 長博

  日本免疫学会総会・学術集会記録, Nov. 2003

  
• 慢性骨髄性白血病(CML)におけるRap1シグナル系の関与

  米谷 耕平; 服部 雅一; 湊 長博

  日本癌学会総会記事, Aug. 2003

  
• SPA-1遺伝子破壊マウスにおける慢性骨髄性白血病の発症

  石田 大輔; 米谷 耕平; 服部 雅一; 湊 長博

  日本癌学会総会記事, Aug. 2003

  
• SPA-1遺伝子破壊マウスにおける慢性骨髄性白血病の発症

  石田 大輔; 米谷 耕平; 服部 雅一; 湊 長博

  日本癌学会総会記事, Oct. 2002

  
• 胸腺内T細胞分化における低分子量G蛋白,Rap1の役割 SPA-1トランスジェニックマウスを用いた解析

  米谷 耕平; 増田 喬子; 河本 宏; 服部 雅一; 湊 長博

  日本免疫学会総会・学術集会記録, Oct. 2002

  
• SPA-1遺伝子破壊マウスにおける免疫不全と慢性骨髄性白血病の発症

  石田 大輔; 米谷 耕平; 服部 雅一; 湊 長博

  日本免疫学会総会・学術集会記録, Oct. 2002

  

Awards

• 2016

  公益財団法人　富山県ひとづくり財団, とやま賞（学術研究部門 医薬分野）
• 2013

  特定非営利活動法人　日本免疫学会, 日本免疫学会　第8回　研究奨励賞

External funds: others

• 京都大学教育研究振興財団　研究活動推進助成

  From 2019, To 31 Mar. 2020

  米谷耕平

Education

Teaching subject(s)

• From 01 Apr. 2024, To 31 Mar. 2025

  ILAS Seminar

  Z001, Spring, Institute for Liberal Arts and Sciences, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  ILAS Seminar

  Z001, Spring, Institute for Liberal Arts and Sciences, 2

Administration

Faculty management (title, position)

• From 01 Apr. 2022, To 31 Mar. 2026

  倫理審査委員会