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Ito, Tsuyoshi

The Kyoto University Museum Assistant Professor

Ito, Tsuyoshi
list
    Last Updated :2024/06/25

    Research

    Research Topics, Overview of the research

    • Research Topics

      Biogeography, evolution, and phylogeny
    • Overview of the research

      1) Hybridization and its consequences in macaques
      2) Demographic history and morphological diversity of Japanese macaques

    Research Areas

    • Informatics, Database science
    • Humanities & social sciences, Museology
    • Life sciences, Evolutionary biology
    • Life sciences, Biodiversity and systematics
    • Life sciences, Ecology and environmental science

    Papers

    • Hybridization and its impact on the ontogenetic allometry of skulls in macaques
      Tsuyoshi Ito; Ryosuke Kimura; Hikaru Wakamori; Mikiko Tanaka; Ayumi Tezuka; Atsushi J Nagano; Yuzuru Hamada; Yoshi Kawamoto
      Evolution, 11 Nov. 2023, Peer-reviewed, Lead author, Corresponding author
    • Analyzing and visualizing morphological features using machine learning techniques and non‐big data: A case study of macaque mandibles
      Takashi Morita; Tsuyoshi Ito; Hiroki Koda; Hikaru Wakamori; Takeshi Nishimura
      American Journal of Biological Anthropology, 13 Jan. 2022, Peer-reviewed
    • Ontogenetic differences in mandibular morphology of two related macaque species and its adaptive implications
      Naoto Toyoda; Tsuyoshi Ito; Tamaki Sato; Takeshi Nishimura
      The Anatomical Record, 25 Apr. 2022, Peer-reviewed
    • Phylogeographic history of Japanese macaques
      Tsuyoshi Ito; Takashi Hayakawa; Nami Suzuki–Hashido; Yuzuru Hamada; Yosuke Kurihara; Goro Hanya; Akihisa Kaneko; Takayoshi Natsume; Seitaro Aisu; Takeaki Honda; Syuji Yachimori; Tomoko Anezaki; Toshinori Omi; Shin‐ichi Hayama; Mikiko Tanaka; Hikaru Wakamori; Hiroo Imai; Yoshi Kawamoto
      Journal of Biogeography, Jun. 2021, Peer-reviewed, Lead author, Corresponding author
    • Stomach and colonic microbiome of wild Japanese macaques
      Wanyi Lee; Takashi Hayakawa; Yosuke Kurihara; Maho Hanzawa; Akiko Sawada; Akihisa Kaneko; Yoshiki Morimitsu; Takayoshi Natsume; Seitaro Aisu; Tsuyoshi Ito; Takeaki Honda; Goro Hanya
      American Journal of Primatology, May 2021, Peer-reviewed
    • Chipping and wear patterns in extant primate and fossil hominin molars: ‘Functional’ cusps are associated with extensive wear but low levels of fracture
      Ian Towle; Carolina Loch; Joel D. Irish; Alessio Veneziano; Tsuyoshi Ito
      Journal of Human Evolution, Feb. 2021, Peer-reviewed
    • Secondary contact and genomic admixture between rhesus and long‐tailed macaques in the Indochina Peninsula
      Tsuyoshi Ito; Sreetharan Kanthaswamy; Srichan Bunlungsup; Robert F. Oldt; Paul Houghton; Yuzuru Hamada; Suchinda Malaivijitnond
      Journal of Evolutionary Biology, 31 Jul. 2020, Peer-reviewed, Lead author, Corresponding author
    • Effects of different segmentation methods on geometric morphometric data collection from primate skulls
      Tsuyoshi Ito
      Methods in Ecology and Evolution, Nov. 2019, Peer-reviewed, Lead author, Corresponding author
    • Shape variation in the facial part of the cranium in macaques and African papionins using geometric morphometrics
      Nishimura T; Morimoto N; Ito T
      Primates, 2019, Peer-reviewed
    • Evidence of different climatic adaptation strategies in humans and non-human primates
      L. T. Buck; I. De Groote; Y. Hamada; B. R. Hassett; T. Ito; J. T. Stock
      Scientific Reports, 30 Jul. 2019, Peer-reviewed
    • Phenotypic plasticity in the mandibular morphology of Japanese macaques: captive–wild comparison
      Kamaluddin SN; Tanaka M; Wakamori H; Nishimura T; Ito T
      Royal Society Open Science, 2019, Peer-reviewed, Last author, Corresponding author
    • Biogeographic variation in skull morphology across the Kra Isthmus in dusky leaf monkeys
      Ito T; Koyabu D
      Journal of Zoological Systematics and Evolutionary Research, Oct. 2018, Peer-reviewed, Lead author, Corresponding author
    • Phylogenetic relationship of a fossil macaque (Macaca cf. robusta) from the Korean Peninsula to extant species of macaques based on zygomaxillary morphology
      Tsuyoshi Ito; Yung-jo Lee; Takeshi D. Nishimura; Mikiko Tanaka; Jong-yoon Woo; Masanaru Takai
      Journal of Human Evolution, 01 Jun. 2018, Peer-reviewed, Lead author, Corresponding author
    • Computed tomography examinations of the surface and internal morphologies of the upper face in Ryukyu Islanders and mainland Japanese population
      Ito T; Kimura R; Ryukoden A; Tsuchiya N; Murayama S; Ishida H
      Anthropological Science, 2018, Peer-reviewed, Lead author, Corresponding author
    • Quantitation of nasal development in the early prenatal period using geometric morphometrics and MRI: a new insight into the critical period of Binder phenotype
      Motoki Katsube; Shigehito Yamada; Reina Miyazaki; Yutaka Yamaguchi; Haruyuki Makishima; Tetsuya Takakuwa; Akira Yamamoto; Yosuke Fujii; Naoki Morimoto; Tsuyoshi Ito; Hirohiko Imai; Shigehiko Suzuki
      Prenatal Diagnosis, Aug. 2017, Peer-reviewed
    • Enigmatic Diversity of the Maxillary Sinus in Macaques and Its Possible Role as a Spatial Compromise in Craniofacial Modifications
      Tsuyoshi Ito; Takeshi D. Nishimura
      Evolutionary Biology, 13 Feb. 2016, Peer-reviewed, Lead author, Corresponding author
    • Phylogenetic inference of Primates Including Extinct Taxa
      Ito T
      Primate Research, Jun. 2016, Peer-reviewed, Lead author, Corresponding author
    • Geographic variation in nasal cavity form among three human groups from the Japanese Archipelago: Ecogeographic and functional implications
      Hitoshi Fukase; Tsuyoshi Ito; Hajime Ishida
      American Journal of Human Biology, 08 Sep. 2015
    • Maxillary sinus variation in hybrid macaques: implications for the genetic basis of craniofacial pneumatization
      Tsuyoshi Ito; Yoshi Kawamoto; Yuzuru Hamada; Takeshi D. Nishimura
      Biological Journal of the Linnean Society, 24 Apr. 2015, Peer-reviewed, Lead author, Corresponding author
    • The source-filter theory of whistle-like calls in marmosets: Acoustic analysis and simulation of helium-modulated voices
      Hiroki Koda; Isao T. Tokuda; Masumi Wakita; Tsuyoshi Ito; Takeshi Nishimura
      The Journal of the Acoustical Society of America, Jun. 2015, Peer-reviewed
    • Contribution of the maxillary sinus to the modularity and variability of nasal cavity shape in Japanese macaques
      Tsuyoshi Ito; Takeshi D. Nishimura; Yuzuru Hamada; Masanaru Takai
      Primates, Jan. 2015, Peer-reviewed, Lead author
    • Aplasia of the maxillary sinus in a Tibetan macaque (Macaca thibetana) with implications for its evolutionary loss and reacquisition
      Takeshi D. Nishimura; Tsuyoshi Ito
      Primates, Oct. 2014, Peer-reviewed
    • Computed tomography examination of the face of Macaca anderssoni (Early Pleistocene, Henan, northern China): Implications for the biogeographic history of Asian macaques
      Tsuyoshi Ito; Takeshi D. Nishimura; Jan Ove R. Ebbestad; Masanaru Takai
      Journal of Human Evolution, Jul. 2014, Peer-reviewed, Lead author, Corresponding author
    • Ecogeographical and phylogenetic effects on craniofacial variation in macaques
      Tsuyoshi Ito; Takeshi Nishimura; Masanaru Takai
      American Journal of Physical Anthropology, May 2014, Peer-reviewed, Lead author, Corresponding author
    • Nasal architecture in Procynocephalus wimani (Early Pleistocene, China) and implications for its phyletic relationship with Paradolichopithecus
      TAKESHI D. NISHIMURA; TSUYOSHI ITO; WATARU YANO; JAN OVE R. EBBESTAD; MASANARU TAKAI
      Anthropological Science, 2014, Peer-reviewed
    • New Hystrix (Mammalia, Rodentia) from the late Miocene/early Pliocene of Myanmar
      Yuichiro Nishioka; Zin-Maung-Maung-Thein; Naoko Egi; Takehisa Tsubamoto; Takeshi Nishimura; Tsuyoshi Ito; Thaung-Htike; Masanaru Takai
      Journal of Vertebrate Paleontology, Jul. 2011, Peer-reviewed
    • Allometry and Interspecific Differences in the Facial Cranium of Two Closely Related Macaque Species
      Tsuyoshi Ito; Takeshi Nishimura; Masanaru Takai
      Anatomy Research International, 25 May 2011, Peer-reviewed, Lead author, Corresponding author
    • Identification of the Tooth Class of Molars in Japanese Macaques (Macaca fuscata) by Using a Geometric Morphometric Method
      Tsuyoshi ITO; Shintaro OGIONO; Yuichiro NISHIOKA; Masanaru TAKAI
      Primate Research, 2010, Peer-reviewed, Lead author, Corresponding author
    • Reappraisal of Macaca speciosa subfossilis from the Late Pleistocene of Northern Vietnam Based on the Analysis of Cranial Anatomy
      Tsuyoshi Ito; Takeshi D. Nishimura; Brigitte Senut; Thomas Koppe; Jacque Treil; Masanaru Takai
      International Journal of Primatology, 24 Jul. 2009, Peer-reviewed, Lead author, Corresponding author

    Misc.

    • 教養部・第三高等学校由来動物標本; 第一部 脊椎動物標本
      本川雅治; 松沼瑞樹; 岡部晋也; 伊藤毅; 西川 完途
      京都大学総合博物館収蔵資料目録, Mar. 2023
    • Introduction of practical examples of outreach activities for high school students at academic conferences: Issue-finding approach through dialogue with researchers
      Takuya MATSUMOTO; Tsuyoshi ITO; Yusuke WATANABE; Rikai SAWAFUJI; Naofumi NAKAGAWA
      Primate Research, 09 Dec. 2022
    • 京都大学ホームカミングデイ—京都大学総合博物館探訪—
      伊藤毅
      京都大学総合博物館ニュースレター, Dec. 2022, Lead author, Corresponding author
    • 金華山のサルの骨格標本に見られる歯の欠損
      若森 参; 伊藤 毅
      宮城県のニホンザル, Aug. 2017

    Presentations

    • マカクザル交雑群のadmixture mappingによる頭蓋形態の種間差に関連する遺伝子多型の探索
      伊藤毅; 川本芳; 濱田穣; 若森参; 手塚あゆみ; 永野惇; 木村亮介
      第122回日本解剖学会大会(2017/04, 長崎), Mar. 2017, The Japanese Association of Anatomists, Invited
    • Genomic admixture and morphological variations in the hybrids between invasive Taiwanese (Macaca cyclopis) and native Japanese macaques (M. fuscata)
      Ito T; Kawamoto Y; Hamada Y; Wakamori H; Tezuka A; Tezuka A; Nagano AJ; Kimura R
      5th Asian Primate Symposium (October 2016, Sri Jayewardenepura), Oct. 2016
    • The genetoc basis of craniofacial variation in hybrid macaques
      Ito T; Kawamoto Y; Hamada Y; Wakamori H; Nagano AJ; Tezuka A; Kimura R
      International Proimatological Society and the American Society of Primatologists (August 2016, Chicago), Aug. 2016, International Primatological Society, Invited
    • 表現型多様性の遺伝的基盤の解明に向けたマカク交雑群のゲノムワイドSNP解析
      伊藤毅; 川本芳; 濱田穣; 若森参; 手塚あゆみ; 永野惇; 木村亮介
      第32回日本霊長類学会大会(2016/07, 鹿児島), Jul. 2016
    • Semi-automatic evaluation of human cranial variations based on computed tomography images
      Tsuyoshi ITO; Ryosuke KIMURA; Azusa RYUKODEN; Hitoshi FUKASE; Nanae TSUCHIYA; Sadayuki MURAYAMA; Hajime ISHIDA
      第69回日本人類学会大会(2015/10, 東京), Oct. 2015
    • 化石から探るマカク属霊長類の歴史生物地理
      伊藤 毅
      第31回日本霊長類学会大会, 19 Jul. 2015, Primate Society of Japan, Invited
    • 霊長類マカク属における鼻腔形状の地理的変異
      伊藤毅; 西村剛; 高井正成
      第29回日本霊長類学会・日本哺乳類学会2013年度合同大会(2013/09,岡山), Sep. 2013
    • マカク属霊長類における顔面頭蓋形状のアロメトリーと地理的多様性
      伊藤毅; 西村剛; 高井正成
      日本霊長類学会第28回大会(2012/07,名古屋), Jul. 2012
    • Paleobiogeography of Macaca (Mammalia: Primates) in East Asia
      Ito T; Nishimura T; Takai M
      Second international symposium on East Asian vertebrate species diversity (July 2012, Kyoto), Jul. 2012
    • 東アジアにおける霊長類マカク属の進化史:頭骨内部構造の変異と系統的意義
      伊藤毅; 西村剛; 高井正成
      日本地球惑星科学連合2012年度連合大会(2012/05,千葉), May 2012
    • Climatic influences on cranial variation in Macaca fascicularis and M. fuscata
      Ito T; Nishimura T; Takai M
      International Primatological Society (September 2010, Kyoto), Sep. 2010
    • A geometric morphometric analysis of a molar shape of Japanese macaque
      Ito T; Ogino S; Takai M
      The 3rd International Symposium of the Biodiversity Global COE Project (July 2009, Kyoto), Jul. 2009
    • ベトナム北部の後期更新世とみられる堆積物から見つかったマカク頭骨化石の形態学的研究
      伊藤毅; 高井正成; 西村剛; Bスニュー
      日本古生物学会2009年年会(2009/06,千葉), Jun. 2009
    • マカク属における鼻腔と上顎洞の形態変異とその系統解析への応用
      伊藤毅; 高井正成; 西村剛; Tコッペ; Bスニュー; Jブラガ; Aベック,Jトレイユ
      日本霊長類学会第24回大会(2008/07,東京), Jul. 2008
    • 隔離分布を示すColopha属アブラムシの進化史
      伊藤毅; 秋元信一
      日本昆虫学会・日本応用動物昆虫学会共催北海道支部大会(2006年度)(2007/01,札幌), Jan. 2007
    • 隔離分布を示すColopha属アブラムシの系統解析
      伊藤毅; 秋元信一
      日本昆虫学会第66回大会(2006/09, 鹿児島), Sep. 2006

    External funds: Kakenhi

    • Phylogeographic history and local adaptation in Japanese macaques: application of population genomics and morphometrics
      Grant-in-Aid for Early-Career Scientists
      Basic Section 45030:Biodiversity and systematics-related
      Kyoto University
      Tsuyoshi Ito
      From 01 Apr. 2019, To 31 Mar. 2021, Project Closed
      系統地理;生態ニッチモデリング;集団遺伝;ゲノム;コンピュータ断層撮影;集団史;進化量的遺伝学;形態
    • アカゲザル種群の進化過程の解明
      Grant-in-Aid for Scientific Research (C)
      Basic Section 45050:Physical anthropology-related
      Kyoto University
      濱田 穣
      From 01 Apr. 2019, To 31 Mar. 2021, Discontinued
      アカゲザル種群;系統地理学;RAD-Seq;集団遺伝学的解析;更新世気候変動;Macaca mulatta;種群;進化;ゲノム解析;生態学的ニッチモデリング
    • Experimental studies of the phonation in non-human primates for elucidating the speech evolution
      Grant-in-Aid for Scientific Research (A)
      Medium-sized Section 45:Biology at organismal to population levels and anthropology, and related fields
      Kyoto University
      西村 剛
      From 01 Apr. 2019, To 31 Mar. 2024, Granted
      霊長類;話しことば;声帯振動;音声;喉頭;声帯膜;進化プロセス;生態振動
    • The process of hybridization and its consequence on phenotypic diversity in primates
      Grant-in-Aid for Young Scientists (B)
      Kyoto University
      Tsuyoshi Ito
      From 01 Apr. 2017, To 31 Mar. 2019, Project Closed
      遺伝子浸透;マカクザル;形態;東南アジア;生殖隔離;交雑帯;RAD-Seq;エコゲノミクス
    • Biological studies of the diversity of phonatory mechanism in non-human primates and of the evolutionary process for human phonation with medical engineering approaches
      Grant-in-Aid for Scientific Research (B)
      Kyoto University
      Takeshi Nishimura
      From 01 Apr. 2016, To 31 Mar. 2020, Project Closed
      人類学;行動学;進化;動物;生物・生体工学;声帯振動;音声生理;サル類;生物・生工学;ヘリウム音声;喉頭;吹鳴実験
    • Population history of the Yakushima Japanese macaques and genetic and morphological consequences of population bottlenecks
      Grant-in-Aid for Scientific Research (C)
      Basic Section 45030:Biodiversity and systematics-related
      Kyoto University
      伊藤 毅
      From 01 Apr. 2023, To 31 Mar. 2027, Granted
      集団史;系統地理;ボトルネック;頭蓋;ほぼ中立説
    list
      Last Updated :2024/06/25

      Education

      Teaching subject(s)

      • From 01 Apr. 2024, To 31 Mar. 2025
        Museum Exhibition: Design and Communication
        W302, Spring, Institute for Liberal Arts and Sciences, 2
      • From 01 Apr. 2024, To 31 Mar. 2025
        Practical on Anatomy and Osteology
        9654, Fall, Graduate School of Science, 1
      • From 01 Apr. 2024, To 31 Mar. 2025
        Advanced Lectures in Primate Systematics and Phylogeny
        9645, Fall, Graduate School of Science, 1
      • From 01 Apr. 2024, To 31 Mar. 2025
        Fundamental Primate Evolution and Phylogeny
        9638, Spring, Graduate School of Science, 1
      • From 01 Apr. 2024, To 31 Mar. 2025
        Seminar on Primate Systematics and Phylogeny D
        9665, Fall, Graduate School of Science, 2
      • From 01 Apr. 2024, To 31 Mar. 2025
        Seminar on Primate Systematics and Phylogeny C
        9664, Spring, Graduate School of Science, 2
      • From 01 Apr. 2024, To 31 Mar. 2025
        Seminar on Primate Systematics and Phylogeny B
        9663, Fall, Graduate School of Science, 2
      • From 01 Apr. 2024, To 31 Mar. 2025
        Seminar on Primate Systematics and Phylogeny A
        9662, Spring, Graduate School of Science, 2
      • From 01 Apr. 2023, To 31 Mar. 2024
        Museum Exhibition: Design and Communication
        W302, Spring, Institute for Liberal Arts and Sciences, 2
      • From 01 Apr. 2023, To 31 Mar. 2024
        Fundamental Primate Evolution and Phylogeny
        9638, Spring, Graduate School of Science, 1
      • From 01 Apr. 2023, To 31 Mar. 2024
        Practical on Anatomy and Osteology
        9654, Fall, Graduate School of Science, 1
      • From 01 Apr. 2023, To 31 Mar. 2024
        Advanced Lectures in Primate Systematics and Phylogeny
        9645, Fall, Graduate School of Science, 1
      • From 01 Apr. 2023, To 31 Mar. 2024
        Seminar on Primate Systematics and Phylogeny A
        9662, Spring, Graduate School of Science, 2
      • From 01 Apr. 2023, To 31 Mar. 2024
        Seminar on Primate Systematics and Phylogeny D
        9665, Fall, Graduate School of Science, 2
      • From 01 Apr. 2023, To 31 Mar. 2024
        Seminar on Primate Systematics and Phylogeny C
        9664, Spring, Graduate School of Science, 2
      • From 01 Apr. 2023, To 31 Mar. 2024
        Seminar on Primate Systematics and Phylogeny B
        9663, Fall, Graduate School of Science, 2
      • From 01 Apr. 2022, To 31 Mar. 2023
        Seminar on Primate Systematics and Phylogeny C
        9664, Spring, Graduate School of Science, 2
      • From 01 Apr. 2022, To 31 Mar. 2023
        Fundamental Primate Evolution and Phylogeny
        9638, Spring, Graduate School of Science, 1
      • From 01 Apr. 2022, To 31 Mar. 2023
        Seminar on Primate Systematics and Phylogeny B
        9663, Fall, Graduate School of Science, 2
      • From 01 Apr. 2022, To 31 Mar. 2023
        Seminar on Primate Systematics and Phylogeny D
        9665, Fall, Graduate School of Science, 2
      • From 01 Apr. 2022, To 31 Mar. 2023
        Advanced Lectures in Primate Systematics and Phylogeny
        9645, Fall, Graduate School of Science, 1
      • From 01 Apr. 2022, To 31 Mar. 2023
        Seminar on Primate Systematics and Phylogeny A
        9662, Spring, Graduate School of Science, 2
      • From 01 Apr. 2022, To 31 Mar. 2023
        Practical on Anatomy and Osteology
        9654, Fall, Graduate School of Science, 1
      • From Apr. 2017, To Mar. 2018
        ILAS Seminar
        Spring, 全学共通科目
      • From Apr. 2017, To Mar. 2018
        Practical on Anatomy and Osteology
        Fall, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Interdisciplinary Seminar on Primatology
        Fall, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Seminar on Primate Evolutionary Morphology A
        Spring, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Seminar on Primate Evolutionary Morphology B
        Fall, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Seminar on Primate Evolutionary Morphology C
        Spring, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Seminar on Primate Evolutionary Morphology D
        Fall, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Advanced Primate Evolutionary Morphology
        Fall, 理学研究科
      • From Apr. 2017, To Mar. 2018
        Fundamental Primate Evolution and Phylogeny
        Spring, 理学研究科
      • From Apr. 2018, To Mar. 2019
        ILAS Seminar
        Spring, 全学共通科目
      • From Apr. 2018, To Mar. 2019
        Practical on Anatomy and Osteology
        Fall, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Seminar on Primate Evolutionary Morphology A
        Spring, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Seminar on Primate Evolutionary Morphology B
        Fall, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Seminar on Primate Evolutionary Morphology C
        Spring, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Seminar on Primate Evolutionary Morphology D
        Fall, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Advanced Primate Evolutionary Morphology
        Fall, 理学研究科
      • From Apr. 2018, To Mar. 2019
        Fundamental Primate Evolution and Phylogeny
        Spring, 理学研究科
      • From Apr. 2019, To Mar. 2020
        ILAS Seminar
        Spring, 全学共通科目
      • From Apr. 2019, To Mar. 2020
        Practical on Anatomy and Osteology
        Fall, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Seminar on Primate Evolutionary Morphology A
        Spring, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Seminar on Primate Evolutionary Morphology B
        Fall, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Seminar on Primate Evolutionary Morphology C
        Spring, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Seminar on Primate Evolutionary Morphology D
        Fall, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Advanced Primate Evolutionary Morphology
        Fall, 理学研究科
      • From Apr. 2019, To Mar. 2020
        Fundamental Primate Evolution and Phylogeny
        Spring, 理学研究科
      • From Apr. 2020, To Mar. 2021
        ILAS Seminar :Evolutionary Morphology of Primates
        Spring, 全学共通科目
      • From Apr. 2020, To Mar. 2021
        Practical on Anatomy and Osteology
        Fall, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Seminar on Primate Evolutionary Morphology A
        Spring, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Seminar on Primate Evolutionary Morphology B
        Fall, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Seminar on Primate Evolutionary Morphology C
        Spring, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Seminar on Primate Evolutionary Morphology D
        Fall, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Advanced Primate Evolutionary Morphology
        Fall, 理学研究科
      • From Apr. 2020, To Mar. 2021
        Fundamental Primate Evolution and Phylogeny
        Spring, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Practical on Anatomy and Osteology
        Fall, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Seminar on Primate Evolutionary Morphology A
        Spring, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Seminar on Primate Evolutionary Morphology B
        Fall, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Seminar on Primate Evolutionary Morphology C
        Spring, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Seminar on Primate Evolutionary Morphology D
        Fall, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Advanced Primate Evolutionary Morphology
        Fall, 理学研究科
      • From Apr. 2021, To Mar. 2022
        Fundamental Primate Evolution and Phylogeny
        Spring, 理学研究科

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