Research

Research Topics, Overview of the research

• Research Topics

  博物館標本を活用した生物多様性情報学的研究
  博物館標本の収集、維持管理、データベース構築、デザイン、展示

Research Interests

• 博物館
• データベース
• 生物多様性情報学
• 生物地理学
• 展示
• デザイン
• バイオインフォマティクス
• natural history
• biodiversity

Research Areas

• Humanities & social sciences, Geography
• Humanities & social sciences, Museology
• Life sciences, Biodiversity and systematics
• Life sciences, Ecology and environmental science
• Life sciences, Genomics
• Life sciences, Systems genomics
• Informatics, Database science
• Informatics, Biological, health, and medical informatics

Papers

• Hybridization and its impact on the ontogenetic allometry of skulls in macaques

  Tsuyoshi Ito; Ryosuke Kimura; Hikaru Wakamori; Mikiko Tanaka; Ayumi Tezuka; Atsushi J Nagano; Yuzuru Hamada; Yoshi Kawamoto

  Evolution, 11 Nov. 2023, Peer-reviewed, Lead author, Corresponding author

  
• Analyzing and visualizing morphological features using machine learning techniques and non‐big data: A case study of macaque mandibles

  Takashi Morita; Tsuyoshi Ito; Hiroki Koda; Hikaru Wakamori; Takeshi Nishimura

  American Journal of Biological Anthropology, 13 Jan. 2022, Peer-reviewed

  
• Ontogenetic differences in mandibular morphology of two related macaque species and its adaptive implications

  Naoto Toyoda; Tsuyoshi Ito; Tamaki Sato; Takeshi Nishimura

  The Anatomical Record, 25 Apr. 2022, Peer-reviewed

  
• Phylogeographic history of Japanese macaques

  Tsuyoshi Ito; Takashi Hayakawa; Nami Suzuki–Hashido; Yuzuru Hamada; Yosuke Kurihara; Goro Hanya; Akihisa Kaneko; Takayoshi Natsume; Seitaro Aisu; Takeaki Honda; Syuji Yachimori; Tomoko Anezaki; Toshinori Omi; Shin‐ichi Hayama; Mikiko Tanaka; Hikaru Wakamori; Hiroo Imai; Yoshi Kawamoto

  Journal of Biogeography, Jun. 2021, Peer-reviewed, Lead author, Corresponding author

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• Stomach and colonic microbiome of wild Japanese macaques

  Wanyi Lee; Takashi Hayakawa; Yosuke Kurihara; Maho Hanzawa; Akiko Sawada; Akihisa Kaneko; Yoshiki Morimitsu; Takayoshi Natsume; Seitaro Aisu; Tsuyoshi Ito; Takeaki Honda; Goro Hanya

  American Journal of Primatology, May 2021, Peer-reviewed

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• Chipping and wear patterns in extant primate and fossil hominin molars: ‘Functional’ cusps are associated with extensive wear but low levels of fracture

  Ian Towle; Carolina Loch; Joel D. Irish; Alessio Veneziano; Tsuyoshi Ito

  Journal of Human Evolution, Feb. 2021, Peer-reviewed

  
• Secondary contact and genomic admixture between rhesus and long‐tailed macaques in the Indochina Peninsula

  Tsuyoshi Ito; Sreetharan Kanthaswamy; Srichan Bunlungsup; Robert F. Oldt; Paul Houghton; Yuzuru Hamada; Suchinda Malaivijitnond

  Journal of Evolutionary Biology, 31 Jul. 2020, Peer-reviewed, Lead author, Corresponding author

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• Effects of different segmentation methods on geometric morphometric data collection from primate skulls

  Tsuyoshi Ito

  Methods in Ecology and Evolution, Nov. 2019, Peer-reviewed, Lead author, Corresponding author

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• Shape variation in the facial part of the cranium in macaques and African papionins using geometric morphometrics

  Nishimura T; Morimoto N; Ito T

  Primates, 2019, Peer-reviewed

  
• Evidence of different climatic adaptation strategies in humans and non-human primates

  L. T. Buck; I. De Groote; Y. Hamada; B. R. Hassett; T. Ito; J. T. Stock

  Scientific Reports, 30 Jul. 2019, Peer-reviewed

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• Phenotypic plasticity in the mandibular morphology of Japanese macaques: captive–wild comparison

  Kamaluddin SN; Tanaka M; Wakamori H; Nishimura T; Ito T

  Royal Society Open Science, 2019, Peer-reviewed, Last author, Corresponding author

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• Biogeographic variation in skull morphology across the Kra Isthmus in dusky leaf monkeys

  Ito T; Koyabu D

  Journal of Zoological Systematics and Evolutionary Research, Oct. 2018, Peer-reviewed, Lead author, Corresponding author

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• Phylogenetic relationship of a fossil macaque (Macaca cf. robusta) from the Korean Peninsula to extant species of macaques based on zygomaxillary morphology

  Tsuyoshi Ito; Yung-jo Lee; Takeshi D. Nishimura; Mikiko Tanaka; Jong-yoon Woo; Masanaru Takai

  Journal of Human Evolution, 01 Jun. 2018, Peer-reviewed, Lead author, Corresponding author

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• Computed tomography examinations of the surface and internal morphologies of the upper face in Ryukyu Islanders and mainland Japanese population

  Ito T; Kimura R; Ryukoden A; Tsuchiya N; Murayama S; Ishida H

  Anthropological Science, 2018, Peer-reviewed, Lead author, Corresponding author

  
• Quantitation of nasal development in the early prenatal period using geometric morphometrics and MRI: a new insight into the critical period of Binder phenotype

  Motoki Katsube; Shigehito Yamada; Reina Miyazaki; Yutaka Yamaguchi; Haruyuki Makishima; Tetsuya Takakuwa; Akira Yamamoto; Yosuke Fujii; Naoki Morimoto; Tsuyoshi Ito; Hirohiko Imai; Shigehiko Suzuki

  Prenatal Diagnosis, Aug. 2017, Peer-reviewed

  
• Enigmatic Diversity of the Maxillary Sinus in Macaques and Its Possible Role as a Spatial Compromise in Craniofacial Modifications

  Tsuyoshi Ito; Takeshi D. Nishimura

  Evolutionary Biology, 13 Feb. 2016, Peer-reviewed, Lead author, Corresponding author

  
• Phylogenetic inference of Primates Including Extinct Taxa

  Ito T

  Primate Research, Jun. 2016, Peer-reviewed, Lead author, Corresponding author

  
• Geographic variation in nasal cavity form among three human groups from the Japanese Archipelago: Ecogeographic and functional implications

  Hitoshi Fukase; Tsuyoshi Ito; Hajime Ishida

  American Journal of Human Biology, 08 Sep. 2015

  
• Maxillary sinus variation in hybrid macaques: implications for the genetic basis of craniofacial pneumatization

  Tsuyoshi Ito; Yoshi Kawamoto; Yuzuru Hamada; Takeshi D. Nishimura

  Biological Journal of the Linnean Society, 24 Apr. 2015, Peer-reviewed, Lead author, Corresponding author

  
• The source-filter theory of whistle-like calls in marmosets: Acoustic analysis and simulation of helium-modulated voices

  Hiroki Koda; Isao T. Tokuda; Masumi Wakita; Tsuyoshi Ito; Takeshi Nishimura

  The Journal of the Acoustical Society of America, Jun. 2015, Peer-reviewed

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• Contribution of the maxillary sinus to the modularity and variability of nasal cavity shape in Japanese macaques

  Tsuyoshi Ito; Takeshi D. Nishimura; Yuzuru Hamada; Masanaru Takai

  Primates, Jan. 2015, Peer-reviewed, Lead author

  
• Aplasia of the maxillary sinus in a Tibetan macaque (Macaca thibetana) with implications for its evolutionary loss and reacquisition

  Takeshi D. Nishimura; Tsuyoshi Ito

  Primates, Oct. 2014, Peer-reviewed

  
• Computed tomography examination of the face of Macaca anderssoni (Early Pleistocene, Henan, northern China): Implications for the biogeographic history of Asian macaques

  Tsuyoshi Ito; Takeshi D. Nishimura; Jan Ove R. Ebbestad; Masanaru Takai

  Journal of Human Evolution, Jul. 2014, Peer-reviewed, Lead author, Corresponding author

  
• Ecogeographical and phylogenetic effects on craniofacial variation in macaques

  Tsuyoshi Ito; Takeshi Nishimura; Masanaru Takai

  American Journal of Physical Anthropology, May 2014, Peer-reviewed, Lead author, Corresponding author

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• Nasal architecture in Procynocephalus wimani (Early Pleistocene, China) and implications for its phyletic relationship with Paradolichopithecus

  TAKESHI D. NISHIMURA; TSUYOSHI ITO; WATARU YANO; JAN OVE R. EBBESTAD; MASANARU TAKAI

  Anthropological Science, 2014, Peer-reviewed

  
• New Hystrix (Mammalia, Rodentia) from the late Miocene/early Pliocene of Myanmar

  Yuichiro Nishioka; Zin-Maung-Maung-Thein; Naoko Egi; Takehisa Tsubamoto; Takeshi Nishimura; Tsuyoshi Ito; Thaung-Htike; Masanaru Takai

  Journal of Vertebrate Paleontology, Jul. 2011, Peer-reviewed

  
• Allometry and Interspecific Differences in the Facial Cranium of Two Closely Related Macaque Species

  Tsuyoshi Ito; Takeshi Nishimura; Masanaru Takai

  Anatomy Research International, 25 May 2011, Peer-reviewed, Lead author, Corresponding author

  
• Identification of the Tooth Class of Molars in Japanese Macaques (Macaca fuscata) by Using a Geometric Morphometric Method

  Tsuyoshi ITO; Shintaro OGIONO; Yuichiro NISHIOKA; Masanaru TAKAI

  Primate Research, 2010, Peer-reviewed, Lead author, Corresponding author

  
• Reappraisal of Macaca speciosa subfossilis from the Late Pleistocene of Northern Vietnam Based on the Analysis of Cranial Anatomy

  Tsuyoshi Ito; Takeshi D. Nishimura; Brigitte Senut; Thomas Koppe; Jacque Treil; Masanaru Takai

  International Journal of Primatology, 24 Jul. 2009, Peer-reviewed, Lead author, Corresponding author

  

Misc.

• 北館2階展示ケースのLED化

  伊藤毅

  京都大学総合博物館ニュースレター, Mar. 2024, Lead author, Corresponding author

  
• 村田泰隆コレクションの内容

  伊藤毅

  京都大学総合博物館ニュースレター, Aug. 2023, Lead author, Corresponding author

  
• 教養部・第三高等学校由来動物標本; 第一部 脊椎動物標本

  本川雅治; 松沼瑞樹; 岡部晋也; 伊藤毅; 西川 完途

  京都大学総合博物館収蔵資料目録, Mar. 2023

  
• Introduction of practical examples of outreach activities for high school students at academic conferences: Issue-finding approach through dialogue with researchers

  Takuya MATSUMOTO; Tsuyoshi ITO; Yusuke WATANABE; Rikai SAWAFUJI; Naofumi NAKAGAWA

  Primate Research, 09 Dec. 2022

  
• 京都大学ホームカミングデイ—京都大学総合博物館探訪—

  伊藤毅

  京都大学総合博物館ニュースレター, Dec. 2022, Lead author, Corresponding author

  
• 金華山のサルの骨格標本に見られる歯の欠損

  若森 参; 伊藤 毅

  宮城県のニホンザル, Aug. 2017

  

Presentations

• 霊長類における種間交雑

  伊藤毅

  京都大学総合博物館セミナー第118回, 17 Jun. 2022

  
• 博物館に長期保存される標本

  第40回日本霊長類学会大会公開シンポジウム「私たちがサルを長く見ていたいワケ」, 14 Jul. 2024, Invited

  
• Butterfly collection in the Kyoto University Museum

  Ito Tsuyoshi

  Kyoto University Museum (KUM) - NTU Museum of Zoology Joint Symposium, 26 Jul. 2023

  
• マカクザル交雑群のadmixture mappingによる頭蓋形態の種間差に関連する遺伝子多型の探索

  伊藤毅; 川本芳; 濱田穣; 若森参; 手塚あゆみ; 永野惇; 木村亮介

  第122回日本解剖学会大会（2017/04, 長崎）, Mar. 2017, The Japanese Association of Anatomists, Invited

  
• Genomic admixture and morphological variations in the hybrids between invasive Taiwanese (Macaca cyclopis) and native Japanese macaques (M. fuscata)

  Ito T; Kawamoto Y; Hamada Y; Wakamori H; Tezuka A; Tezuka A; Nagano AJ; Kimura R

  5th Asian Primate Symposium (October 2016, Sri Jayewardenepura), Oct. 2016

  
• The genetoc basis of craniofacial variation in hybrid macaques

  Ito T; Kawamoto Y; Hamada Y; Wakamori H; Nagano AJ; Tezuka A; Kimura R

  International Proimatological Society and the American Society of Primatologists (August 2016, Chicago), Aug. 2016, International Primatological Society, Invited

  
• 表現型多様性の遺伝的基盤の解明に向けたマカク交雑群のゲノムワイドSNP解析

  伊藤毅; 川本芳; 濱田穣; 若森参; 手塚あゆみ; 永野惇; 木村亮介

  第32回日本霊長類学会大会（2016/07, 鹿児島）, Jul. 2016

  
• Semi-automatic evaluation of human cranial variations based on computed tomography images

  Tsuyoshi ITO; Ryosuke KIMURA; Azusa RYUKODEN; Hitoshi FUKASE; Nanae TSUCHIYA; Sadayuki MURAYAMA; Hajime ISHIDA

  第69回日本人類学会大会（2015/10, 東京）, Oct. 2015

  
• 化石から探るマカク属霊長類の歴史生物地理

  伊藤 毅

  第31回日本霊長類学会大会, 19 Jul. 2015, Primate Society of Japan, Invited

  
• 霊長類マカク属における鼻腔形状の地理的変異

  伊藤毅; 西村剛; 高井正成

  第29回日本霊長類学会・日本哺乳類学会2013年度合同大会（2013/09，岡山）, Sep. 2013

  
• マカク属霊長類における顔面頭蓋形状のアロメトリーと地理的多様性

  伊藤毅; 西村剛; 高井正成

  日本霊長類学会第28回大会（2012/07，名古屋）, Jul. 2012

  
• Paleobiogeography of Macaca (Mammalia: Primates) in East Asia

  Ito T; Nishimura T; Takai M

  Second international symposium on East Asian vertebrate species diversity (July 2012, Kyoto), Jul. 2012

  
• 東アジアにおける霊長類マカク属の進化史：頭骨内部構造の変異と系統的意義

  伊藤毅; 西村剛; 高井正成

  日本地球惑星科学連合2012年度連合大会（2012/05，千葉）, May 2012

  
• Climatic influences on cranial variation in Macaca fascicularis and M. fuscata

  Ito T; Nishimura T; Takai M

  International Primatological Society (September 2010, Kyoto), Sep. 2010

  
• A geometric morphometric analysis of a molar shape of Japanese macaque

  Ito T; Ogino S; Takai M

  The 3rd International Symposium of the Biodiversity Global COE Project (July 2009, Kyoto), Jul. 2009

  
• ベトナム北部の後期更新世とみられる堆積物から見つかったマカク頭骨化石の形態学的研究

  伊藤毅; 高井正成; 西村剛; Bスニュー

  日本古生物学会2009年年会（2009/06，千葉）, Jun. 2009

  
• マカク属における鼻腔と上顎洞の形態変異とその系統解析への応用

  伊藤毅; 高井正成; 西村剛; Tコッペ; Bスニュー; Jブラガ; Aベック,Jトレイユ

  日本霊長類学会第24回大会（2008/07，東京）, Jul. 2008

  
• 隔離分布を示すColopha属アブラムシの進化史

  伊藤毅; 秋元信一

  日本昆虫学会・日本応用動物昆虫学会共催北海道支部大会（2006年度）（2007/01，札幌）, Jan. 2007

  
• 隔離分布を示すColopha属アブラムシの系統解析

  伊藤毅; 秋元信一

  日本昆虫学会第66回大会（2006/09, 鹿児島）, Sep. 2006

  

Books and Other Publications

• 動物・植物・鉱物から探る古代中国

  浅見真生; 伊藤毅; 孫国平; 王永磊; 高井正成, Contributor, 中国浙江省の考古遺跡から出土したサルの化石—東アジアにおけるマカクザルの分布域の変化に関する考察—

  六一書房, Mar. 2024

  Find Kyoto University's collection of books :

  
• 霊長類学の百科事典 = The encyclopedia of primatology

  伊藤毅, Contributor, 霊長類の頭蓋

  丸善出版, Jul. 2023

  Find Kyoto University's collection of books :

  
• 霊長類学の百科事典 = The encyclopedia of primatology

  伊藤毅, Contributor, ニホンザルの起源と進化

  丸善出版, Jul. 2023

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• 日本のサル学のあした : 霊長類研究という「人間学」の可能性

  中川, 尚史; 友永, 雅己; 山極, 寿一, Contributor, マカク属霊長類の進化史を化石から辿る

  京都通信社, Dec. 2012

  Find Kyoto University's collection of books :

  
• 新しい霊長類学 : 人を深く知るための100問100答

  京都大学霊長類研究所, Contributor

  講談社, Sep. 2009

  Find Kyoto University's collection of books :

  

Works

• diploid2haploid

  From 25 May 2020, To Present

  
• sam_sexing

  From 24 May 2020

  
• 解剖学・骨学実習〜計測・解析編2〜

  伊藤毅

  From 17 Dec. 2021, To Present

  
• 解剖学・骨学実習—計測・解析編—

  伊藤毅

  From 14 Dec. 2020, To Present

  

External funds: Kakenhi

• Phylogeographic history and local adaptation in Japanese macaques: application of population genomics and morphometrics

  Grant-in-Aid for Early-Career Scientists

  Basic Section 45030:Biodiversity and systematics-related

  Kyoto University

  Tsuyoshi Ito

  From 01 Apr. 2019, To 31 Mar. 2021, Project Closed

  系統地理;生態ニッチモデリング;集団遺伝;ゲノム;コンピュータ断層撮影;集団史;進化量的遺伝学;形態

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19K16211/
• アカゲザル種群の進化過程の解明

  Grant-in-Aid for Scientific Research (C)

  Basic Section 45050:Physical anthropology-related

  Kyoto University

  濱田 穣

  From 01 Apr. 2019, To 31 Mar. 2021, Discontinued

  アカゲザル種群;系統地理学;RAD-Seq;集団遺伝学的解析;更新世気候変動;Macaca mulatta;種群;進化;ゲノム解析;生態学的ニッチモデリング

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19K06865/
• Experimental studies of the phonation in non-human primates for elucidating the speech evolution

  Grant-in-Aid for Scientific Research (A)

  Medium-sized Section 45:Biology at organismal to population levels and anthropology, and related fields

  Kyoto University

  西村 剛

  From 01 Apr. 2019, To 31 Mar. 2024, Granted

  霊長類;話しことば;声帯振動;音声;喉頭;声帯膜;進化プロセス;生態振動

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19H01002/
• The process of hybridization and its consequence on phenotypic diversity in primates

  Grant-in-Aid for Young Scientists (B)

  Kyoto University

  Tsuyoshi Ito

  From 01 Apr. 2017, To 31 Mar. 2019, Project Closed

  遺伝子浸透;マカクザル;形態;東南アジア;生殖隔離;交雑帯;RAD-Seq;エコゲノミクス

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-17K15195/
• Biological studies of the diversity of phonatory mechanism in non-human primates and of the evolutionary process for human phonation with medical engineering approaches

  Grant-in-Aid for Scientific Research (B)

  Kyoto University

  Takeshi Nishimura

  From 01 Apr. 2016, To 31 Mar. 2020, Project Closed

  人類学;行動学;進化;動物;生物・生体工学;声帯振動;音声生理;サル類;生物・生工学;ヘリウム音声;喉頭;吹鳴実験

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-16H04848/
• Population history of the Yakushima Japanese macaques and genetic and morphological consequences of population bottlenecks

  Grant-in-Aid for Scientific Research (C)

  Basic Section 45030:Biodiversity and systematics-related

  Kyoto University

  伊藤 毅

  From 01 Apr. 2023, To 31 Mar. 2027, Granted

  集団史;系統地理;ボトルネック;頭蓋;ほぼ中立説

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23K05905/

Education

Teaching subject(s)

• From 01 Apr. 2024, To 31 Mar. 2025

  Museum Exhibition: Design and Communication

  W302, Spring, Institute for Liberal Arts and Sciences, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Practical on Anatomy and Osteology

  9654, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Advanced Lectures in Primate Systematics and Phylogeny

  9645, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Fundamental Primate Evolution and Phylogeny

  9638, Spring, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Primate Systematics and Phylogeny D

  9665, Fall, Graduate School of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Primate Systematics and Phylogeny C

  9664, Spring, Graduate School of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Primate Systematics and Phylogeny B

  9663, Fall, Graduate School of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Primate Systematics and Phylogeny A

  9662, Spring, Graduate School of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Museum Exhibition: Design and Communication

  W302, Spring, Institute for Liberal Arts and Sciences, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Fundamental Primate Evolution and Phylogeny

  9638, Spring, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Practical on Anatomy and Osteology

  9654, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Advanced Lectures in Primate Systematics and Phylogeny

  9645, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Primate Systematics and Phylogeny A

  9662, Spring, Graduate School of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Primate Systematics and Phylogeny D

  9665, Fall, Graduate School of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Primate Systematics and Phylogeny C

  9664, Spring, Graduate School of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Primate Systematics and Phylogeny B

  9663, Fall, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Primate Systematics and Phylogeny C

  9664, Spring, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Fundamental Primate Evolution and Phylogeny

  9638, Spring, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Primate Systematics and Phylogeny B

  9663, Fall, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Primate Systematics and Phylogeny D

  9665, Fall, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Advanced Lectures in Primate Systematics and Phylogeny

  9645, Fall, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Primate Systematics and Phylogeny A

  9662, Spring, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Practical on Anatomy and Osteology

  9654, Fall, Graduate School of Science, 1
• From Apr. 2017, To Mar. 2018

  ILAS Seminar

  Spring, 全学共通科目
• From Apr. 2017, To Mar. 2018

  Practical on Anatomy and Osteology

  Fall, 理学研究科
• From Apr. 2017, To Mar. 2018

  Interdisciplinary Seminar on Primatology

  Fall, 理学研究科
• From Apr. 2017, To Mar. 2018

  Seminar on Primate Evolutionary Morphology A

  Spring, 理学研究科
• From Apr. 2017, To Mar. 2018

  Seminar on Primate Evolutionary Morphology B

  Fall, 理学研究科
• From Apr. 2017, To Mar. 2018

  Seminar on Primate Evolutionary Morphology C

  Spring, 理学研究科
• From Apr. 2017, To Mar. 2018

  Seminar on Primate Evolutionary Morphology D

  Fall, 理学研究科
• From Apr. 2017, To Mar. 2018

  Advanced Primate Evolutionary Morphology

  Fall, 理学研究科
• From Apr. 2017, To Mar. 2018

  Fundamental Primate Evolution and Phylogeny

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  ILAS Seminar

  Spring, 全学共通科目
• From Apr. 2018, To Mar. 2019

  Practical on Anatomy and Osteology

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Primate Evolutionary Morphology A

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Primate Evolutionary Morphology B

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Primate Evolutionary Morphology C

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Primate Evolutionary Morphology D

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Advanced Primate Evolutionary Morphology

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Fundamental Primate Evolution and Phylogeny

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  ILAS Seminar

  Spring, 全学共通科目
• From Apr. 2019, To Mar. 2020

  Practical on Anatomy and Osteology

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Primate Evolutionary Morphology A

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Primate Evolutionary Morphology B

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Primate Evolutionary Morphology C

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Primate Evolutionary Morphology D

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Advanced Primate Evolutionary Morphology

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Fundamental Primate Evolution and Phylogeny

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  ILAS Seminar :Evolutionary Morphology of Primates

  Spring, 全学共通科目
• From Apr. 2020, To Mar. 2021

  Practical on Anatomy and Osteology

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Primate Evolutionary Morphology A

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Primate Evolutionary Morphology B

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Primate Evolutionary Morphology C

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Primate Evolutionary Morphology D

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Advanced Primate Evolutionary Morphology

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Fundamental Primate Evolution and Phylogeny

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Practical on Anatomy and Osteology

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Primate Evolutionary Morphology A

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Primate Evolutionary Morphology B

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Primate Evolutionary Morphology C

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Primate Evolutionary Morphology D

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Advanced Primate Evolutionary Morphology

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Fundamental Primate Evolution and Phylogeny

  Spring, 理学研究科

Academic, Social Contribution

Academic Contribution

• 京都大学総合博物館2023年度特別展「蝶に会える日—村田泰隆コレクション展— Vol.2 東南アジアが育んだ多様性」

  Planning, management, etc.

  京都大学総合博物館, From 29 Nov. 2023, To 24 Dec. 2023

  
• 京都大学総合博物館2022年度特別展「蝶に会える日—村田泰隆コレクション展— Vol.1 アゲハチョウ類の多様性」

  Planning, management, etc.

  京都大学総合博物館, 京都大学総合博物館, From 08 Feb. 2023, To 05 Mar. 2023

  

Social Contribution

• ニホンザルの歯牙年齢推定

  Official expert, Investigator

  From 30 Aug. 2023, To 31 Aug. 2023

  
• 蝶の多様性

  Lecturer

  京都大学総合博物館, 京都大学総合博物館, From 13 Feb. 2023, To 13 Feb. 2023

  
• 混血がもたらすサルの多様性

  Lecturer

  日本人類学会・日本霊長類学会連合大会公開コロキウム—ヒト屋/サル屋と高校生の対面議論—, From 18 Sep. 2022, To 18 Sep. 2022