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Macpherson, Tom

Graduate School of Pharmaceutical Sciences, Division of Bioinformatics and Chemical Genomics Senior Lecturer/ Junior Associate Professor

Macpherson, Tom
list
    Last Updated :2026/10/30

    Basic Information

    Faculty

    • Institute for Liberal Arts and Sciences (ILAS)

    Email Address

    • Email Address

      macpherson.tom.3jkyoto-u.ac.jp

    Professional Memberships

    • From 2016, To Present
      Society for Neuroscience
    • From May 2014, To Present
      The Physiological Society of Japan
    • From May 2014, To Present
      The Japan Neuroscience Society

    Academic Degree

    • 17 Feb. 2014
      Ph.D.(Sussex University)

    Academic Resume (Undergraduate School/Majors)

    • The College of Richard Collyer, 卒業
    • Sussex University, School of Psychology, Psychology, 卒業

    Research History

    • From Apr. 2025, To Present
      Kyoto University, Faculty of Pharmaceutical Sciences, Senior Lecturer
    • From 2020, To May 2025
      Osaka University, Institute for Protein Research, Assistant Professor
    • From 2018, To 2020
      Osaka University, Institute for Protein Research, Specially Appointed Assistant Professor
    • From 2017, To 2018
      Kyoto University, Graduate School of Medicine, Postdoctoral Researcher
    • From 2014, To 2017
      Kyoto University, Graduate School of Medicine, JSPS Research Fellow
    • From 2014, To 2014
      Sussex University, Neuroscience, Postdoctoral Researcher

    ID,URL

    researchmap URL

    list
      Last Updated :2026/10/30

      Research

      Research Interests

      • Optogenetics
      • Mouse
      • Prefrontal Cortex
      • GABA
      • Dopamine
      • Calcium Imaging
      • Fiber Photometry
      • Single-cell level In-vivo Brain Imaging
      • Striatum
      • Nucleus Accumbens
      • Classical Conditioning
      • Psychiatric Conditions
      • Cognitive Flexibility
      • Decision-making
      • Learning
      • Behavioral Neuroscience
      • Neuroscience

      Research Areas

      • Life sciences, Animals: biochemistry, physiology, behavioral science
      • Life sciences, Psychiatry
      • Life sciences, Neuroscience - general

      Papers

      • The slow-acting G-protein G z defines the duration of circadian rest time
        Youichi Tanaka; Yuto Fujita; Tom Macpherson; Aoi Harui; Sumihiro Kunisue; Genzui Setsu; Xinyan Shao; Daisuke Ono; Takatoshi Hikida; Yoshihisa Fujita; Michiyuki Matsuda; Hitoshi Okamura; Emi Hasegawa; Masao Doi
        27 May 2026
      • Inside insight: decoding how insight emerges from competing world models
        Kengo Inutsuka; Tadaaki Nishioka; Tom Macpherson; Mana Fujiwara; Takatoshi Hikida; Honda Naoki
        26 May 2026
      • Essential Roles of Heparan Sulfate Endosulfatase Sulf1 in Reward and Aversion Learning Through Distinct Dopamine D1 and D2 Receptor Pathways in Male Mice.
        Ken Miya; Kent Ohta; Kazuko Keino-Masu; Takuya Okada; Seiya Mizuno; Satoru Takahashi; Tom Macpherson; Takatoshi Hikida; Masayuki Masu
        Journal of neurochemistry, Jan. 2026, Peer-reviewed
      • Flexible value coding in the mesolimbic dopamine system depending on internal water and sodium balance.
        Takaaki Ozawa; Issei Nakagawa; Yuuki Uchida; Mayuka Abe; Tom Macpherson; Yuichi Yamashita; Takatoshi Hikida
        NPJ science of food, 30 Sep. 2025, Peer-reviewed
      • 味覚情報伝達の分子・神経機構 味覚の相乗作用と情動生成を司るトップダウン神経回路
        小澤 貴明; 尾山 賀信; 柴田 智弘; 阿部 万友佳; 後藤 健太朗; 米丸 ひなの; 松本 悠真; 岩本 涼太郎; 櫻井 航輝; マクファーソン・トム; 疋田 貴俊
        Journal of Oral Biosciences Supplement, Nov. 2024
      • Sex-dependent differences in the ability of nicotine to modulate discrimination learning and cognitive flexibility in mice.
        Yoshiatsu Aomine; Yuto Shimo; Koki Sakurai; Mayuka Abe; Tom Macpherson; Takaaki Ozawa; Takatoshi Hikida
        Journal of neurochemistry, 17 Sep. 2024, Peer-reviewed
      • Clozapine N-oxide, compound 21, and JHU37160 do not influence effortful reward-seeking behavior in mice
        Yoshiatsu Aomine; Yoshinobu Oyama; Koki Sakurai; Tom Macpherson; Takaaki Ozawa; Takatoshi Hikida
        Psychopharmacology, Oct. 2023, Peer-reviewed
      • α4-containing GABAAreceptors on DRD2-neurons of the nucleus accumbens mediate instrumental responding for conditioned reinforcers, and its potentiation by cocaine
        Tom Macpherson; Claire I. Dixon; Jonathan Robertson; Marsha M. Sindarto; Patricia H. Janak; Delia Belelli; Jeremy J. Lambert; David N. Stephens; Sarah L. King
        eNeuro, 08 Aug. 2023, Peer-reviewed, Lead author, Corresponding author
      • Dual roles for nucleus accumbens core dopamine D1-expressing neurons projecting to the substantia nigra pars reticulata in limbic and motor control in male mice
        Suthinee Attachaipanich; Takaaki Ozawa; Tom Macpherson; Takatoshi Hikida
        eNeuro, 26 May 2023, Peer-reviewed, Last author, Corresponding author
      • Error-related signaling in nucleus accumbens D2 receptor-expressing neurons guides inhibition-based choice behavior in mice
        Tadaaki Nishioka; Suthinee Attachaipanich; Kosuke Hamaguchi; Michael Lazarus; Alban de Kerchove d’Exaerde; Tom Macpherson; Takatoshi Hikida
        Nature Communications, 21 Apr. 2023, Peer-reviewed, Last author, Corresponding author
      • Circuit, molecular, and developmental mechanisms in decision-making behavior
        Tom Macpherson; Minae Niwa; Hirofumi Morishita; Takatoshi Hikida
        Frontiers in Neuroscience, Apr. 2023, Peer-reviewed, Lead author, Corresponding author
      • Elucidating the role of neurotransmission via D1 and D2 dopamine receptors in the mechanisms of motor control, learning, and memory
        Nae Saito; Makoto Itakura; Kazuki Tainaka; Tom Macpherson; Takatoshi Hikida; Shun Yamaguchi; Asako Sato; Tadashi Okubo; Satomi Chiken; Atsushi Nambu; Toshikuni Sasaoka
        Japanese Journal of Biological Psychiatry, Sep. 2022, Peer-reviewed
      • Importin α3 (KPNA3) Deficiency Augments Effortful Reward-Seeking Behavior in Mice
        Yoshiatsu Aomine; Koki Sakurai; Tom Macpherson; Takaaki Ozawa; Yoichi Miyamoto; Yoshihiro Yoneda; Masahiro Oka; Takatoshi Hikida
        Frontiers in Neuroscience, 30 Jun. 2022, Peer-reviewed
      • Nucleus Accumbens Core Dopamine D2 Receptor-Expressing Neurons Control Reversal Learning but Not Set-Shifting in Behavioral Flexibility in Male Mice
        Tom Macpherson; Ji Yoon Kim; Takatoshi Hikida
        Frontiers in Neuroscience, 28 Jun. 2022, Peer-reviewed, Lead author, Corresponding author
      • Error-related Signaling in Nucleus Accumbens D2 Receptor-expressing Neurons Guides Avoidance-based Goal-directed Behavior
        Tadaaki Nishioka; Tom Macpherson; Kosuke Hamaguchi; Takatoshi Hikida
        Research Square, 05 Jan. 2022, Last author, Corresponding author
      • Natural and Artificial Intelligence: A brief introduction to the interplay between AI and neuroscience research.
        Tom Macpherson; Anne Churchland; Terry Sejnowski; James DiCarlo; Yukiyasu Kamitani; Hidehiko Takahashi; Takatoshi Hikida
        Neural networks : the official journal of the International Neural Network Society, Dec. 2021, Peer-reviewed, Lead author
      • Parallel and hierarchical neural mechanisms for adaptive and predictive behavioral control
        Tom Macpherson; Masayuki Matsumoto; Hiroaki Gomi; Jun Morimoto; Eiji Uchibe; Takatoshi Hikida
        Neural Networks, Sep. 2021, Peer-reviewed, Lead author
      • Distinct Roles of Dopamine D1 and D2 Receptor-expressing Neurons in the Nucleus Accumbens for a Strategy Dependent Decision Making
        Tadaaki Nishioka; Tom Macpherson; Kosuke Hamaguchi; Takatoshi Hikida
        BioRxiv, 05 Aug. 2021
      • Effects of Importin α1/KPNA1 deletion and adolescent social isolation stress on psychiatric disorder-associated behaviors in mice.
        Koki Sakurai; Taichi Itou; Makiko Morita; Emiko Kasahara; Tetsuji Moriyama; Tom Macpherson; Takaaki Ozawa; Yoichi Miyamoto; Yoshihiro Yoneda; Atsuo Sekiyama; Masahiro Oka; Takatoshi Hikida
        PloS one, 2021, Peer-reviewed
      • Neurotransmission through dopamine D1 receptors is required for aversive memory formation and Arc activation in the cerebral cortex.
        Nae Saito; Kazuki Tainaka; Tom Macpherson; Takatoshi Hikida; Shun Yamaguchi; Toshikuni Sasaoka
        Neuroscience research, 05 May 2020, Peer-reviewed
      • Nucleus accumbens pathways control cell-specific gene expression in the medial prefrontal cortex.
        Takatoshi Hikida; Shuhei Yao; Tom Macpherson; Ayumi Fukakusa; Makiko Morita; Haruhide Kimura; Keisuke Hirai; Tatsuya Ando; Hiroyoshi Toyoshiba; Akira Sawa
        Scientific reports, 04 Feb. 2020, Peer-reviewed, Lead author
      • Adolescent psychosocial stress enhances sensitization to cocaine exposure in genetically vulnerable mice.
        Takatoshi Hikida; Makiko Morita; Mahomi Kuroiwa; Tom Macpherson; Takahide Shuto; Naoki Sotogaku; Minae Niwa; Akira Sawa; Akinori Nishi
        Neuroscience research, Feb. 2020, Peer-reviewed
      • Role of basal ganglia neurocircuitry in the pathology of psychiatric disorders.
        Tom Macpherson; Takatoshi Hikida
        Psychiatry and clinical neurosciences, Jun. 2019, Peer-reviewed, Lead author
      • Preproenkephalin-expressing ventral pallidal neurons control inhibitory avoidance learning.
        Tom Macpherson; Hiroyuki Mizoguchi; Akihiro Yamanaka; Takatoshi Hikida
        Neurochemistry international, Jun. 2019, Peer-reviewed, Lead author
      • Response to 'Mood and Affect'.
        Tom Macpherson; Takatoshi Hikida
        Psychiatry and clinical neurosciences, Jun. 2019, Peer-reviewed, Lead author
      • Nucleus Accumbens Dopamine D1-Receptor-Expressing Neurons Control the Acquisition of Sign-Tracking to Conditioned Cues in Mice.
        Tom Macpherson; Takatoshi Hikida
        Frontiers in neuroscience, 2018, Peer-reviewed, Lead author
      • [Basal Ganglia Circuit Mechanisms in Cognitive Learning].
        Takatoshi Hikida; Tom MacPherson; Makiko Morita
        Nihon shinkei seishin yakurigaku zasshi = Japanese journal of psychopharmacology, Apr. 2017, Peer-reviewed
      • Neural mechanisms of the nucleus accumbens circuit in reward and aversive learning.
        Takatoshi Hikida; Makiko Morita; Tom Macpherson
        Neuroscience research, Jul. 2016, Peer-reviewed, Last author
      • Nucleus accumbens dopamine D2-receptor expressing neurons control behavioral flexibility in a place discrimination task in the IntelliCage.
        Tom Macpherson; Makiko Morita; Yanyan Wang; Toshikuni Sasaoka; Akira Sawa; Takatoshi Hikida
        Learning & memory (Cold Spring Harbor, N.Y.), Jul. 2016, Peer-reviewed, Lead author
      • Tonic inhibition of accumbal spiny neurons by extrasynaptic α4βδ GABAA receptors modulates the actions of psychostimulants.
        Edward P Maguire; Tom Macpherson; Jerome D Swinny; Claire I Dixon; Murray B Herd; Delia Belelli; David N Stephens; Sarah L King; Jeremy J Lambert
        The Journal of neuroscience : the official journal of the Society for Neuroscience, 15 Jan. 2014, Peer-reviewed, Lead author
      • Striatal direct and indirect pathways control decision-making behavior.
        Tom Macpherson; Makiko Morita; Takatoshi Hikida
        Frontiers in psychology, 2014, Peer-reviewed, Lead author
      • Seeing with sound? exploring different characteristics of a visual-to-auditory sensory substitution device.
        David Brown; Tom Macpherson; Jamie Ward
        Perception, 2011, Peer-reviewed

      Misc.

      • 味覚の相乗作用と情動生成を司るトップダウン神経回路
        小澤貴明; 尾山賀信; 柴田智弘; 阿部万友佳; 後藤健太朗; 米丸ひなの; 松本悠真; 岩本涼太郎; 櫻井航輝; マクファーソン トム; 疋田貴俊
        Journal of Oral Biosciences Supplement (Web), 2024
      • 腕傍核から腹側被蓋野への神経回路の光遺伝学を用いた活性化によるマウスの場所嗜好性並びに回避行動に与える影響
        尾山賀信; 小澤貴明; マクファーソン トム; 疋田貴俊
        日本生物学的精神医学会(Web), 2024
      • 恐怖条件づけ中のマウス脳内における領域特異的なドーパミン放出動態
        小澤貴明; 梅本和宏; 岩本涼太郎; 中村萌; 松本悠真; 尾山賀信; 柴田智弘; MACPHERSON Tom; 疋田貴俊
        日本生物学的精神医学会(Web), 2024
      • アルコール依存症関連行動を制御する核-細胞質間輸送メカニズム
        青峰良淳; 青峰良淳; 櫻井航輝; 櫻井航輝; マクファーソン トム; マクファーソン トム; 小澤貴明; 小澤貴明; 宮本洋一; 米田悦啓; 岡正啓; 疋田貴俊; 疋田貴俊
        日本生物学的精神医学会(Web), 2024
      • アルコール消費および耐性を上昇させる核-細胞質間輸送異常
        青峰良淳; 櫻井航輝; 足立淳; マクファーソン トム; 小澤貴明; 宮本洋一; 米田悦啓; 岡正啓; 疋田貴俊
        日本分子生物学会年会プログラム・要旨集(Web), 2024
      • The role of dopaminergic circuits on the enhancing effects of umami on salt seeking behavior in mice.
        柴田智弘; 米丸ひなの; 松本悠真; 中村萌; 岩本涼太郎; 尾山賀信; 櫻井航輝; MACPHERSON Tom; 疋田貴俊; 小澤貴明
        日本味と匂学会誌, 2021
      • Nucleus Accumbens D1 Receptor-Expressing Neurons Control Autoshaping Behavior
        Tom Macpherson; Takatoshi Hikida
        NEUROPSYCHOPHARMACOLOGY, Nov. 2017
      • Social Isolated DISC1 Mutant Mice Displayed High Sensitivity to Chronic Cocaine Exposure and Rolipram Treatment
        Takatoshi Hikida; Makiko Morita; Mahomi Kuroiwa; Tom Macpherson; Taichi Itou; Takahide Shuto; Naoki Sotogaku; Minae Niwa; Akira Sawa; Akinori Nishi
        NEUROPSYCHOPHARMACOLOGY, Dec. 2015
      • 社会的ストレス下においてDISC1変異マウスのコカイン感受性はPDE4により制御される(Cocaine-induced behavioral sensitization is regulated by PDE4 in social stressed DISC1 mutant mice)
        疋田 貴俊; 森田 真規子; マクファーソン・トム; 伊藤 大一; 丹羽 美苗; 黒岩 真帆美; 澤 明; 西 昭徳
        日本生物学的精神医学会・日本神経精神薬理学会合同年会プログラム・抄録集, Sep. 2015

      Presentations

      • Striatal control of cue-guided decision-making and its impairment in psychiatric conditions
        Tom Macpherson
        47th Annual Meeting of the Japan Neuroscience Society (JNS), 26 Jul. 2024, Invited
      • Striatal mechanisms guiding reward, aversion, and decision-making
        Tom Macpherson
        Inscopix Webinar, 20 Jun. 2024, Invited
      • Error-related signaling in nucleus accumbens D2 receptor-expressing neurons guides inhibition-based choice behavior
        Tom Macpherson
        Molecular and Cellular Cognition Society (MCCS) Meeting, 09 Nov. 2023, Invited
      • Striatal mechanisms of discrimination learning and their dysfunction in psychiatric conditions
        Tom Macpherson
        5th Annual Meeting of the UK-Japan Neuroscience Sympsoium, Sep. 2023, Invited
      • Striatal mechanisms of cue-guided decision-making and its dysfunction in schizophrenia model mice
        Tom Macpherson
        46th Annual Meeting of the Japan Neuroscience Society (JNS), Jul. 2023, Invited
      • Cell-type-specific control of reward and aversive signaling in the Nucleus Accumbens
        Tom Macpherson; Takatoshi Hikida
        100th Annual Meeting of The Physiological Society of Japan Meeting, 14 Mar. 2023, Invited
      • Striatal mechanisms of discrimination learning and their dysfunction in schizophrenia model mice
        Macpherson T; Morita M; Sawa A; Hikida T
        52nd Annual Meeting of the Society for Neuroscience (SfN), 16 Nov. 2022, Invited
      • Neural mechanisms in reward and aversion signaling
        Macpherson T; Attachaipanich S; Nishioka T; Ozawa T; Hikida T
        National Institute of Physiological Science (NIPS) Emotion Conference, 08 Sep. 2022, Invited
      • Neural mechanisms in reward and aversion signaling
        Macpherson T; Attachaipanich S; Nishioka T; Ozawa T; Hikida T
        45th Annual Meeting of the Japan Neuroscience Society (JNS), 30 Jun. 2022, Invited
      • Parallel Neural Circuit Mechanisms in Behavioral Flexibility
        Tom Macpherson
        15th Annual Meeting of the Japanese Society for Motor Control, Oct. 2021, Invited
      • Nucleus accumbens D2-receptor-expressing neurons regulate reversal learning in the Attentional Set-Shifting Test
        Tom Macpherson; Jiyoon Kim; Takatoshi Hikida
        44th Annual Meeting of the Japan Neuroscience Society (JNS), Jun. 2021
      • Nucleus accumbens pathways control cell-specific gene expression in the medial prefrontal cortex
        Tom Macpherson; Shuhei Yao; Ayumi Fukakusa; Makiko Morita; Haruhide Kimura; Keisuke Hirai; Tatsuya Ando; Hiroshi Toyoshiba; Akira Sawa
        43rd Annual Meeting of the Japan Neuroscience Society (JNS), Jun. 2020
      • A role for Enkephalin-expressing ventral pallidal neurons in controlling aversive Pavlovian Conditioning
        Tom Macpherson
        97th Annual Meeting of the Physiological Society of Japan, Mar. 2020, Invited
      • Altered medial prefrontal cortex gene expression following nucleus accumbens pathway neurotransmission blocking
        Tom Macpherson; Shuhei Yao; Ayumi Fukakusa; Makiko Morita; Haruhide Kimura; Keisuke Hirai; Tatsuya Ando; Hiroshi Toyoshiba; Akira Sawa
        10th Takeda Science Foundation Symposium, Jan. 2020
      • A role for nucleus accumbens D1-/D2-neuron pathways in controlling learning impaired in psychiatric disorders
        Tom Macpherson
        42nd Annual Meeting of the Japan Neuroscience Society (JNS), Jul. 2019, Invited
      • Ventral pallidum neurons control aversive learning
        Tom Macpherson; Hiroyuki Mizoguchi; Akihiro Yamanaka; Takatoshi Hikida
        48th Annual Meeting of the Society for Neuroscience (SfN), Nov. 2018
      • Ventral pallidum neurons control aversive learning
        Tom Macpherson; Hiroyuki Mizoguchi; Akihiro Yamanaka; Takatoshi Hikida
        11th Meeting of the Federation for European Neuroscience Society (FENS), Jul. 2018
      • Ventral pallidum neurons control aversive learning
        Tom Macpherson; Hiroyuki Mizoguchi; Akihiro Yamanaka; Takatoshi Hikida
        41st Annual Meeting of the Japan Neuroscience Society (JNS), Jul. 2018
      • Dopamine D2L receptors control behavioral flexibility
        Tom Macpherson
        2nd Joint Symposium on Protein Structure and Function, Dec. 2017, Invited
      • Nucleus Accumbens D1-receptor-expressing neurons control autoshaping behavior
        Tom Macpherson; Takatoshi Hikida
        47th Annual Meeting of the Society for Neuroscience (SfN), Nov. 2017
      • Nucleus Accumbens D1-receptor-expressing neurons control attribution of incentive salience in an autoshaping task
        Tom Macpherson
        17th Biennial Meeting of the European Behavioral Pharmacology Society (EBPS), Aug. 2017
      • Nucleus Accumbens D1-receptor-expressing neurons control autoshaping behavior
        Tom Macpherson; Takatoshi Hikida
        40th Annual Meeting of the Japan Neuroscience Society (JNS), Jul. 2017
      • Nucleus accumbens dopamine D1-receptor-expressing neurons control incentive salience to reward-predictive cues.
        Tom Macpherson
        26th Annual Meeting of the International Behavioral Neuroscience Society (IBNS), Jun. 2017, Invited
      • Neurotransmission in dopamine D2-receptor-expressing nucleus accumbens neurons is necessary for behavioral flexibility in an IntelliCage place discrimination task.
        Tom Macpherson
        46th Annual Meeting of the Society for Neuroscience (SfN), Nov. 2016, Invited
      • Nucleus accumbens dopamine D2-receptor expressing neurons control behavioral flexibility in a place learning task
        Tom Macpherson, Makiko Morita, Yanyan Wang, Toshikuni Sasaoka, Akira Sawa, Takatoshi Hikida
        39th Annual Meeting of the Japan Neuroscience Society (JNS), Jul. 2016
      • Nucleus accumbens dopamine D2-receptor expressing neurons control behavioral flexibility in a place learning task in the IntelliCage
        Tom Macpherson, Makiko Morita, Yanyan Wang, Toshikuni Sasaoka, Akira Sawa, Takatoshi Hikida
        10th Meeting of Federation for European Neuroscience Society (FENS), Jul. 2016

      Awards

      • 22 Nov. 2023
        大阪大学賞(若手教員部門)

      External funds: Kakenhi

      • Elucidating the neural mechanisms that support flexible decision-making
        Grant-in-Aid for Scientific Research (C)
        Basic Section 51020:Cognitive and brain science-related
        Kyoto University
        マクファーソン トム
        From 01 Apr. 2026, To 31 Mar. 2029, Granted
        Decision-making;In-vivo imaging;Aging;Cognitive decline
      • 失敗から学習するための神経回路・分子機構とその破綻
        Grant-in-Aid for Scientific Research (B)
        Basic Section 51020:Cognitive and brain science-related
        The University of Osaka
        疋田 貴俊
        From 01 Apr. 2025, To 31 Mar. 2028, Granted
        精神疾患;大脳基底核;ドーパミン;神経回路
      • 戦略的な意思決定の神経機構と精神疾患病態の解明
        Grant-in-Aid for Scientific Research (B)
        Basic Section 51020:Cognitive and brain science-related
        Osaka University
        疋田 貴俊
        From 01 Apr. 2024, To 31 Mar. 2026, Granted
        神経回路;大脳基底核;ドーパミン;精神疾患
      • Investigation of nucleus accumbens subpopulations controlling reward and aversive learning
        Grant-in-Aid for Early-Career Scientists
        Basic Section 46030:Function of nervous system-related
        Osaka University
        Tom Macpherson
        From 01 Apr. 2021, To 31 Mar. 2023, Project Closed
        Reward;Aversion;Learning;Nucleus Accumbens;Basal Ganglia;In-vivo imaging;Optogenetics;In-Vivo Imaging;Behavior
      • Neuronal circuit mechanisms of reward/goal-directed behavior
        Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)
        Complex systems
        Osaka University;Kyoto University
        Takatoshi HIKIDA
        From 30 Jun. 2016, To 31 Mar. 2021, Project Closed
        大脳基底核;神経回路;意思決定;強化学習;神経科学;脳・神経;脳神経疾患;トランスレーショナルリサーチ;生理学;大脳皮質;腹側淡蒼球;ドーパミン受容体;行動柔軟性
      list
        Last Updated :2026/10/30

        Education

        Teaching subject(s)

        • From 01 Apr. 2026, To 31 Mar. 2027
          Basic Biology and Metabolism-E2
          N941, Fall, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2026, To 31 Mar. 2027
          Introduction to Biosciences-E2
          N940, Fall, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2026, To 31 Mar. 2027
          Introduction to Molecular Cell Biology-E2
          N932, Spring, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2026, To 31 Mar. 2027
          Theory and Practice in Scientific Writing and Discussion (Pharmaceutical Sciences, English)A-E3
          W224, Spring, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2025, To 31 Mar. 2026
          Theory and Practice in Scientific Writing and Discussion (Pharmaceutical Sciences, English)A-E3
          W224, Spring, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2025, To 31 Mar. 2026
          Basic Biology and Metabolism-E2
          N941, Fall, Institute for Liberal Arts and Sciences, 2
        • From 01 Apr. 2025, To 31 Mar. 2026
          Introduction to Biosciences-E2
          N940, Fall, Institute for Liberal Arts and Sciences, 2

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