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Terakawa, Tsuyoshi
Graduate School of Science, Division of Biological Sciences Associate Professor
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Last Updated :2025/06/02
Basic Information
Faculty
Professional Memberships
- From Jan. 2023, To PresentThe Molecular Biology Society of Japan
- From Apr. 2010, To PresentThe Biophysical Society of Japan
Academic Degree
Research History
- From Feb. 2024, To PresentKyoto University, Department of Biophysics, Graduate School of Science, Associate Professor
- From Apr. 2018, To Jan. 2024Kyoto University, Department of Biophysics, Graduate School of Science, Assistant Professor
- From Apr. 2017, To Mar. 2018Columbia University, Department of Biochemistry and Molecular Biophysics, Post-Doctoral Research Fellow of the Uehara Memorial Foundation
- From Apr. 2015, To Mar. 2017Columbia University, Department of Biophysics, Postdoctoral Fellow of JSPS
- From Apr. 2014, To Mar. 2015Kyoto University, Department of Biophysics, Postdoctoral Fellow of JSPS (PD)
- From Apr. 2013, To Mar. 2014Kyoto University, Department of Biophysics, Postdoctoral Fellow of JSPS (DC2)
Profile
Profile
In our group, we tackle fundamental questions of life, such as gene expression and DNA replication in eukaryotes, as well as the chromatin structural changes that regulate these processes. Our unique approach combines world-leading single-molecule imaging techniques with large-scale molecular simulations using supercomputers. By integrating experiments, simulations, and machine learning, we aim to elucidate the molecular-level dynamics underlying essential chromatin-based processes such as transcription, replication, and repair, and to contribute to a physicochemical understanding of epigenetics and cell fate regulation. We are always welcoming new members. If you are passionate about understanding life at the molecular level or wish to acquire cutting-edge experimental and computational skills, we encourage you to join us — regardless of your background in life sciences, physics, chemistry, or information science. If you are interested, please feel free to contact us by email (terakawa at biophys.kyoto-u.ac.jp).
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ORCID ID
J-Global ID
Researcher ID
GHW-0050-2022Researcher number
20809652External link
researchmap URL
list
Last Updated :2025/06/02
Research
Research Interests
- Biophysics
- Single molecule fluorescent microscopy
- Nanopore sequencing
- Molecular dynamics simulation
- DNA replication
- DNA repair
- DNA transcription
- Chromatin structure
Research Areas
Papers
- Mechanistic models of asymmetric hand-over-hand translocation and nucleosome navigation by CMG helicaseFritzh Nagae; Yutaka Murata; Masataka Yamauchi; Shoji Takada; Tsuyoshi TerakawabioRxiv, 05 May 2025, Last author, Corresponding author
- Solution AFM imaging and coarse-grained molecular modeling of yeast condensin structural variation coupled to the ATP hydrolysis cycleHiroki Koide; Noriyuki Kodera; Shoji Takada; Tsuyoshi TerakawaJournal of Molecular Biology, 03 May 2025, Peer-reviewed, Last author, Corresponding author
- Minimal protein list for chromatin replication-coupled symmetric histone recycling revealed through in vitro reconstitutionFritz Nagae; Shizuko Endo; Yasuto Murayama; Tsuyoshi TerakawabioRxiv, 19 Nov. 2024, Last author, Corresponding author
- SMC complex unidirectionally translocates DNA by coupling segment capture with an asymmetric kleisin pathMasataka Yamauchi; Giovanni B. Brandani; Tsuyoshi Terakawa; Shoji TakadabioRxiv, 02 May 2024
- Molecular dynamics simulations of human cohesin subunits identify DNA binding sites and their potential roles in DNA loop extrusionChenyang Gu; Shoji Takada; Giovanni B. Brandani; Tsuyoshi TerakawaPLOS Computational Biology, 04 Apr. 2025, Peer-reviewed, Last author, Corresponding author
- Molecular mechanism of parental H3/H4 recycling at a replication forkFritz Nagae; Yasuto Murayama; Tsuyoshi TerakawaNature Communications, 02 Nov. 2024, Peer-reviewed, Last author, Corresponding author
- Histone chaperone Nap1 dismantles an H2A/H2B dimer from a partially unwrapped nucleosomeFritz Nagae; Shoji Takada; Tsuyoshi TerakawaNucleic Acids Research, 13 May 2023, Peer-reviewed, Last author, Corresponding author
- ALS mutations in the TIA-1 prion-like domain trigger highly condensed pathogenic structuresNaotaka Sekiyama; Kiyofumi Takaba; Saori Maki-Yonekura; Ken-ichi Akagi; Yasuko Ohtani; Kayo Imamura; Tsuyoshi Terakawa; Keitaro Yamashita; Daigo Inaoka; Koji Yonekura; Takashi S. Kodama; Hidehito TochioProceedings of the National Academy of Sciences, 16 Sep. 2022, Peer-reviewed
- Coarse-grained molecular dynamics simulations of base-pair mismatch recognition protein MutS sliding along DNAKeisuke Inoue; Shoji Takada; Tsuyoshi TerakawaBiophysics and Physicobiology, 2022, Peer-reviewed, Last author, Corresponding author
- Single-molecule junction spontaneously restored by DNA zipperTakanori Harashima; Shintaro Fujii; Yuki Jono; Tsuyoshi Terakawa; Noriyuki Kurita; Satoshi Kaneko; Manabu Kiguchi; Tomoaki NishinoNature Communications, 01 Oct. 2021, Peer-reviewed
- The lane-switch mechanism for nucleosome repositioning by DNA translocaseFritz Nagae; Giovanni B Brandani; Shoji Takada; Tsuyoshi TerakawaNucleic Acids Research, 07 Aug. 2021, Peer-reviewed, Last author, Corresponding author
- Modeling of DNA binding to the condensin hinge domain using molecular dynamics simulations guided by atomic force microscopy.Hiroki Koide; Noriyuki Kodera; Shveta Bisht; Shoji Takada; Tsuyoshi TerakawaPLOS computational biology, Jul. 2021, Peer-reviewed, Last author, Corresponding author
- Defining the influence of Rad51 and Dmc1 lineage-specific amino acids on genetic recombination.Justin B Steinfeld; Ondrej Beláň; Youngho Kwon; Tsuyoshi Terakawa; Amr Al-Zain; Michael J Smith; J Brooks Crickard; Zhi Qi; Weixing Zhao; Rodney Rothstein; Lorraine S Symington; Patrick Sung; Simon J Boulton; Eric C GreeneGenes & development, 01 Sep. 2019, Peer-reviewed
- Nucleosome Crowding in Chromatin Slows the Diffusion but Can Promote Target Search of Proteins.Ryo Kanada; Tsuyoshi Terakawa; Hiroo Kenzaki; Shoji TakadaBiophysical Journal, 18 Jun. 2019, Peer-reviewed
- The condensin complex is a mechanochemical motor that translocates along DNATsuyoshi Terakawa; Shveta Bisht; Jorine M. Eeftens; Cees Dekker; Christian H. Haering; Eric C. GreeneScience, Nov. 2017, Peer-reviewed, Lead author
- Sequential eviction of crowded nucleoprotein complexes by the exonuclease RecBCD molecular motorTsuyoshi Terakawa; Sy Redding; Timothy D. Silverstein; Eric C. GreeneProceedings of the National Academy of Sciences, Aug. 2017, Peer-reviewed, Lead author
- How one-dimensional diffusion of transcription factors are affected by obstacles: coarse-grained molecular dynamics studyMami Saito; Tsuyoshi Terakawa; Shoji TakadaMolecular Simulation, 2017, Peer-reviewed
- Dynamic Coupling among Protein Binding, Sliding, and DNA Bending Revealed by Molecular DynamicsCheng Tan; Tsuyoshi Terakawa; Shoji TakadaJournal of the American Chemical Society, Jul. 2016, Peer-reviewed
- Modeling Structural Dynamics of Biomolecular Complexes by Coarse-Grained Molecular SimulationsShoji Takada; Ryo Kanada; Cheng Tan; Tsuyoshi Terakawa; Wenfei Li; Hiroo KenzakiAccounts of Chemical Research, Dec. 2015, Peer-reviewed
- p53 dynamics upon response element recognition explored by molecular simulationsTsuyoshi Terakawa; Shoji TakadaScientific Reports, Nov. 2015, Peer-reviewed, Lead author
- Base triplet stepping by the Rad51/RecA family of recombinasesJa Yil Lee; Tsuyoshi Terakawa; Zhi Qi; Justin B. Steinfeld; Sy Redding; YoungHo Kwon; William A. Gaines; Weixing Zhao; Patrick Sung; Eric C. GreeneScience, Aug. 2015, Peer-reviewed
- Multi-scale Ensemble Modeling of Modular Proteins with Intrinsically Disordered Linker Regions: Application to p53Tsuyoshi Terakawa; Junichi Higo; Shoji TakadaBiophysical Journal, Aug. 2014, Peer-reviewed, Lead author
- RESPAC: Method to Determine Partial Charges in Coarse-Grained Protein Model and Its Application to DNA-Binding ProteinsTsuyoshi Terakawa; Shoji TakadaJournal of Chemical Theory and Computation, Feb. 2014, Peer-reviewed, Lead author
- Energy landscape and multiroute folding of topologically complex proteins adenylate kinase and 2ouf-knotWenfei Li; Tsuyoshi Terakawa; Wei Wang; Shoji TakadaProceedings of the National Academy of Sciences, Oct. 2012, Peer-reviewed
- p53 Searches on DNA by Rotation-Uncoupled Sliding at C-Terminal Tails and Restricted Hopping of Core DomainsTsuyoshi Terakawa; Hiroo Kenzaki; Shoji TakadaJournal of the American Chemical Society, Sep. 2012, Peer-reviewed, Lead author
- Multiscale Ensemble Modeling of Intrinsically Disordered Proteins: p53 N-Terminal DomainTsuyoshi Terakawa; Shoji TakadaBiophysical Journal, Sep. 2011, Peer-reviewed, Lead author
- On Easy Implementation of a Variant of the Replica Exchange with Solute Tempering in GROMACSTsuyoshi Terakawa; Tomoshi Kameda; Shoji TakadaJournal of Computational Chemistry, May 2011, Peer-reviewed, Lead author
Misc.
- コンデンシンの分子モーター活性を同定寺川 剛生物物理, Apr. 2018, Peer-reviewed, Invited
- 染色体構造の形成をつかさどる分子モーター：コンデンシン寺川 剛実験医学, Feb. 2018, Invited
- 海外だより ～もし素人がアメリカで実験をはじめたら～寺川 剛生物物理, Mar. 2017, Invited
- コンデンシン複合体はDNAに沿って移動する メカノケミカルモーターである寺川 剛サイエンス誌に載った日本人研究者, 2017, Invited
- リコンビナーゼRad51/RecAファミリーによる 塩基トリプレット歩進寺川 剛サイエンス誌に載った日本人研究者, 2015, Invited
Presentations
- Molecular Dynamics of Histone Recycling in Chromatin ReplicationTsuyoshi TerakawaThe 47th Annual Meeting of the Molecular Biology Society of Japn, 27 Nov. 2024, Invited
- Reconstitution of chromatin replication-coupled symmetric histone recyclingFritz Nagae; Shizuko Endo; Yasuto Murayama; Tsuyoshi Terakawa5th East Asian Single-Molecule Biophysics Symposium, 08 Nov. 2024, Invited
- Biophysical approaches to the molecular mechanisms of histone recyclingTsuyoshi Terakawa第97回日本生化学会大会, 06 Nov. 2024, Invited
- Unveiling the molecular mechanism of histone recycling through biophysical approachesTsuyoshi Terakawa21st IUPAB Congress 2024, 27 Jun. 2024, Invited
- Exploration of the molecular mechanism of histone recycling through molecular dynamics simulations, nanopore sequencing, and biochemical experimentsTsuyoshi Terakawa68th Annual Meeting of the Biophysical Society, Feb. 2024
- Biophysical approaches reveal molecular mechanism of histone recyclingTsuyoshi TerakawaThe 46th Annual Meeting of the Molecular Biology Society of Japan, Dec. 2023, Invited
- Condensin’s journey on DNA with obstaclesTsuyoshi Terakawa4th East Asian Single-Molecule Biophysics Symposium, Sep. 2023, Invited
- トランスロケースとの衝突に伴うヌクレオソームの運命: エピゲノム継承の解明に向けて寺川 剛第15回エピジェネティクス研究会年会, May 2022, Invited
- Nucleosome repositioning dynamics upon collision with a translocaseTsuyoshi TerakawaThe 44th Annual Meeting of the Molecular Biology Society of Japan, Dec. 2021, Invited
- DNA curtain assay of epigenome inheritance at single molecule level寺川 剛さきがけCREST「ゲノム合成」2020年度 合同領域会議, Jan. 2021, Invited
- Reconstitution of protein functions on chromatin curtain寺川 剛第43回分子生物学会年会ワークショップ「Reconstruction of the nucleus - Hierarchical understanding of the functional nucleus by reconstitutional approaches -」, Dec. 2020, Invited
- Toward the molecular mechanism of condensin translocation on DNA寺川 剛新学術領域「発動分子科学」第4回領域会議, Nov. 2020, Invited
- Histone modification and chromatin condensation explored by DNA curtain寺川 剛新学術領域「クロマチン潜在能」領域会議, May 2020, Invited
- DNA curtain assays and molecular dynamics simulations to elucidate the molecular mechanism of DNA transactions寺川 剛2nd ExCELLS Retreat for Young Scientists, Feb. 2020, Invited
- Condensinモーターの発動機構の解明に向けて寺川 剛新学術領域「発動分子科学」第2回領域会議, Jan. 2020
- Single-molecule observation of chromatin condensation induced by Condensin molecular motor寺川 剛第42回日本分子生物学会年会, Dec. 2019, Invited
- Single-molecule DNA curtain assays of epigenetic inheritance寺川 剛第２回キックオフCREST/さきがけ（ゲノム合成領域）合同領域会議, Nov. 2019, Invited
- DNAカーテン法を用いたDNA上の混み合い問題へのアプローチ寺川 剛OCU先端光科学シンポジウム, Oct. 2019, Invited
- DNA curtains: high-throughput single molecule imaging for DNA transactions寺川 剛第57回日本生物物理学会年会, Sep. 2019, Invited
- Condensin is a mechanochemical molecular motor that translocates along DNA寺川 剛理研エピジェネティクスセミナー, Sep. 2019, Invited
- A new type of DNA molecular motor: CondensinTsuyoshi Terakawa2nd East Asian Single-Molecule Biophysics Symposium, Jul. 2019, Invited
- Relationship between histone modification and chromatin condensation explored by DNA curtain assayTsuyoshi Terakawa新学術領域「クロマチン潜在能」 第2回領域会議, Jun. 2019, Invited
- 金・銀ナノ粒子を用いたコンデンシン分子モーターの超高分解能DNAカーテン測定寺川 剛新学術領域「発動分子科学」第2回領域会議, May 2019, Invited
- 染色体凝集の分子基盤: コンデンシン分子モーターの分子観察寺川 剛国立遺伝学研究所 研究集会「ゲノムの維持継承を支える分子基盤の包括的理解とその発展」, Feb. 2019, Invited
- DNA上の新しい分子モーター: コンデンシン寺川 剛分子モーター討論会, Nov. 2018, Invited
- The condensin complex is a mechanochemical molecular motorTsuyoshi TerakawaThe 56th Annual Meeting of The Biophysical Society of Japan, Sep. 2018, Invited
- The condensin complex is a new type of DNA molecular motorTsuyoshi TerakawaThe 79th Okazaki Conference, Sep. 2018, Invited
- The condensin complex is a mechanochemical motor that translocates along DNATsuyoshi TerakawaGenome Integrity Discussion Group (NYAS), Oct. 2017
External funds: Kakenhi
- クロマチンカーテン法によるクロマチン凝集の1分子蛍光顕微鏡観察Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)Biological SciencesKyoto University寺川 剛From 01 Apr. 2021, To 31 Mar. 2023, Project Closedコンデンシン;分子モーター;染色体ループ形成機構;ヌクレオソーム;DNAカーテン法;一分子蛍光イメージング;蛍光顕微鏡観察;クロマチンカーテン;染色体凝集
- 金銀・ナノ粒子を用いたコンデンシン分子モーターの超高分解能DNAカーテン測定Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)Complex systemsKyoto University寺川 剛From 01 Apr. 2019, To 31 Mar. 2021, Project Closedコンデンシン;蛍光顕微鏡観察;DNAカーテン;一分子実験;量子ドット;金ナノ粒子;銀ナノ粒子;1分子蛍光顕微鏡観察;ステップサイズ測定
- DNAカーテン測定によるヒストン化学修飾がクロマチン凝集に与える影響の解明Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)Biological SciencesKyoto University寺川 剛From 01 Apr. 2019, To 31 Mar. 2021, Project Closedコンデンシン;ヌクレオソーム;蛍光顕微鏡観察;DNAカーテン;一分子実験;染色体;ヒストン化学修飾;1分子蛍光顕微鏡観察;染色体凝集;クロマチン;エピジェネティクス
- Molecular mechanisms of a Condensin molecular motor explored by DNA curtain assays and coarse-grained molecular dynamics simulationsGrant-in-Aid for Scientific Research (B)Basic Section 43040:Biophysics-relatedKyoto UniversityTsuyoshi TerakawaFrom 01 Apr. 2019, To 31 Mar. 2022, Project Closed１分子蛍光イメージング;DNAカーテン;粗視化分子動力学シミュレーション;コンデンシン;蛍光顕微鏡観察;一分子実験;分子モーター
- Elucidation of the molecular mechanism of chromatin folding by single molecule experimentsGrant-in-Aid for Research Activity Start-up0701:Biology at molecular to cellular levels, and related fieldsKyoto University寺川 剛From 24 Aug. 2018, To 31 Mar. 2020, Project ClosedDNAカーテン;蛍光顕微鏡;コンデンシン;染色体凝縮;分子モーター;染色体;ヌクレオソーム;全反射蛍光顕微鏡
- ナノポア解析と分子シミュレーションで探るヒストンリサイクリングの鎖バイアス機構Grant-in-Aid for Transformative Research Areas (B)Transformative Research Areas, Section (III)Kyoto University寺川 剛From 01 Apr. 2024, To 31 Mar. 2027, Grantedゲノム複製;エピゲノム継承;ヒストンリサイクリング;ナノポアシーケンシング;分子動力学シミュレーション
- エピゲノム継承への分子・細胞スケールブリッジアプローチの推進Grant-in-Aid for Transformative Research Areas (B)Transformative Research Areas, Section (III)Kyoto University寺川 剛From 01 Apr. 2024, To 31 Mar. 2027, Grantedヒストンリサイクリング;分子動力学シミュレーション;次世代シーケンシング;光ピンセット;染色体再構成
- Molecule- and cell-scale bridging approach to epigenome inheritanceGrant-in-Aid for Transformative Research Areas (B)From 01 Apr. 2024, To 31 Mar. 2027
- 細胞内複雑環境下の相分離による生体分子凝縮体形成の生物物理Grant-in-Aid for Scientific Research (B)Basic Section 43040:Biophysics-relatedKyoto University高田 彰二From 01 Apr. 2024, To 31 Mar. 2027, Granted生体分子凝縮体;液液相分離;転写制御;シナプス可塑性
- Molecule- and cell-scale bridging approach to epigenome inheritanceGrant-in-Aid for Transformative Research Areas (B)From 01 Apr. 2024, To 31 Mar. 2027
list
Last Updated :2025/06/02
Education
Teaching subject(s)
- From 01 Apr. 2025, To 31 Mar. 2026Advanced Course on Biological Science A9011, Spring, Graduate School of Science, 2
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IIC9027, Spring, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IIB9026, Fall, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IIA9025, Spring, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IID9028, Fall, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IA9021, Spring, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics ID9024, Fall, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IC9023, Spring, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Seminar on Theoretical Biophysics IB9022, Fall, Graduate School of Science, 1
- From 01 Apr. 2024, To 31 Mar. 2025Special Study CourseII(Biological Sciences) 238726, Year-long, Faculty of Science, 12
- From 01 Apr. 2024, To 31 Mar. 2025Laboratory work in Biological Science A3762, Spring, Faculty of Science, 2
- From 01 Apr. 2024, To 31 Mar. 2025Biomolecular Science I2716, Spring, Faculty of Science, 2
- From 01 Apr. 2024, To 31 Mar. 2025Biomolecular Science II2715, Fall, Faculty of Science, 2
- From 01 Apr. 2024, To 31 Mar. 2025Seminars in Biological Science B4732, Fall, Faculty of Science, 2
- From 01 Apr. 2024, To 31 Mar. 2025Advanced Course on Biological Science A9011, Spring, Graduate School of Science, 2
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IID9028, Fall, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IIC9027, Spring, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IIB9026, Fall, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IB9022, Fall, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IA9021, Spring, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IIA9025, Spring, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics ID9024, Fall, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Seminar on Theoretical Biophysics IC9023, Spring, Graduate School of Science, 1
- From 01 Apr. 2023, To 31 Mar. 2024Special Study CourseII(Biological Sciences) 238726, Year-long, Faculty of Science, 12
- From 01 Apr. 2023, To 31 Mar. 2024Structural Biology8736, Fall, Faculty of Science, 2
- From 01 Apr. 2023, To 31 Mar. 2024Seminars in Biological Science B4732, Fall, Faculty of Science, 2
- From 01 Apr. 2023, To 31 Mar. 2024Laboratory work in Biological Science B3763, Spring, Faculty of Science, 2
- From 01 Apr. 2023, To 31 Mar. 2024Advanced Course on Biological Science B9012, Fall, Graduate School of Science, 2
- From 01 Apr. 2022, To 31 Mar. 2023Practice of Basic Informatics (Faculty of Science)T003, Fall, Institute for Liberal Arts and Sciences, 2
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IIB9026, Fall, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics ID9024, Fall, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IID9028, Fall, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IC9023, Spring, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Advanced Course on Biological Science B9012, Fall, Graduate School of Science, 2
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IB9022, Fall, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Special Study CourseII(Biological Sciences) 238726, Year-long, Faculty of Science, 12
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IIC9027, Spring, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IIA9025, Spring, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Structural Biology8736, Fall, Faculty of Science, 2
- From 01 Apr. 2022, To 31 Mar. 2023Seminar on Theoretical Biophysics IA9021, Spring, Graduate School of Science, 1
- From 01 Apr. 2022, To 31 Mar. 2023Laboratory work in Biological Science B3763, Spring, Faculty of Science, 2
- From 01 Apr. 2022, To 31 Mar. 2023Seminars in Biological Science B4732, Fall, Faculty of Science, 2
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IASpring, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IBFall, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics ICSpring, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IDFall, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IIASpring, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IIBFall, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IICSpring, 理学研究科
- From Apr. 2018, To Mar. 2019Seminar on Theoretical Biophysics IIDFall, 理学研究科
- From Apr. 2018, To Mar. 2019Seminars in Biological Science BFall, 理学部
- From Apr. 2018, To Mar. 2019Laboratory work in Biological Science EFall, 理学部
- From Apr. 2018, To Mar. 2019Special Study CourseII(Biological Sciences) 23Year-long, 理学部
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IASpring, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IBFall, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics ICSpring, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IDFall, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IIASpring, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IIBFall, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IICSpring, 理学研究科
- From Apr. 2019, To Mar. 2020Seminar on Theoretical Biophysics IIDFall, 理学研究科
- From Apr. 2019, To Mar. 2020Seminars in Biological Science BFall, 理学部
- From Apr. 2019, To Mar. 2020Laboratory work in Biological Science EFall, 理学部
- From Apr. 2019, To Mar. 2020Special Study CourseII(Biological Sciences) 23Year-long, 理学部
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IASpring, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IBFall, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics ICSpring, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IDFall, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IIASpring, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IIBFall, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IICSpring, 理学研究科
- From Apr. 2020, To Mar. 2021Seminar on Theoretical Biophysics IIDFall, 理学研究科
- From Apr. 2020, To Mar. 2021Seminars in Biological Science BFall, 理学部
- From Apr. 2020, To Mar. 2021Laboratory Work in Biological Science EFall, 理学部
- From Apr. 2020, To Mar. 2021Special Study CourseII(Biological Sciences) 23Year-long, 理学部
- From Apr. 2021, To Mar. 2022Structural BiologyFall, 理学部
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IASpring, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IBFall, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics ICSpring, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IDFall, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IIASpring, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IIBFall, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IICSpring, 理学研究科
- From Apr. 2021, To Mar. 2022Seminar on Theoretical Biophysics IIDFall, 理学研究科
- From Apr. 2021, To Mar. 2022Seminars in Biological Science BFall, 理学部
- From Apr. 2021, To Mar. 2022Laboratory Work in Biological Science EFall, 理学部
- From Apr. 2021, To Mar. 2022Advanced Course on Biological Science BFall, 理学研究科
- From Apr. 2021, To Mar. 2022Special Study CourseII(Biological Sciences) 23Year-long, 理学部
Participation in PhD Defense
list
Last Updated :2025/06/02
Academic, Social Contribution
Committee Memberships
Academic Contribution
- Nature CommunicationsPeer review
- 第63回日本生物物理学会年会Planning, management, etc.From 24 Sep. 2025, To 26 Sep. 2025
- IUPAB 2024 (21st IUPAB & 62nd BSJ joint congress)Planning, management, etc.From 24 Jun. 2024, To 28 Jun. 2024
- Frontiers in Molecular BiosciencesPeer review
- Computational and Structural Biotechnology JournalPeer review
- Nature Structural & Molecular BiologyPeer review
- Nucleic Acids ResearchPeer review
- Science AdvancesPeer review
- Biophysical JournalPeer review
