Basic Information

Faculty

• 理学部

Professional Memberships

• From Jan. 2023, To Present

  The Molecular Biology Society of Japan
• From Apr. 2010, To Present

  The Biophysical Society of Japan

Academic Degree

• 修士（理学）(京都大学)
• 博士（理学）(京都大学)

Research History

• From Feb. 2024, To Present

  Kyoto University, Department of Biophysics, Graduate School of Science, Associate Professor
• From Apr. 2018, To Jan. 2024

  Kyoto University, Department of Biophysics, Graduate School of Science, Assistant Professor
• From Apr. 2017, To Mar. 2018

  Columbia University, Department of Biochemistry and Molecular Biophysics, Post-Doctoral Research Fellow of the Uehara Memorial Foundation
• From Apr. 2015, To Mar. 2017

  Columbia University, Department of Biophysics, Postdoctoral Fellow of JSPS
• From Apr. 2014, To Mar. 2015

  Kyoto University, Department of Biophysics, Postdoctoral Fellow of JSPS (PD)
• From Apr. 2013, To Mar. 2014

  Kyoto University, Department of Biophysics, Postdoctoral Fellow of JSPS (DC2)

ID,URL

• J-Global ID

  201801015264093169

• Researcher number

  20809652

• External link

  https://theory.biophys.kyoto-u.ac.jp/terakawa-en

researchmap URL

• https://researchmap.jp/ttterakawa

Research

Research Interests

• Biophysics
• Single molecule fluorescent microscopy
• Nanopore sequencing
• Molecular dynamics simulation
• DNA replication
• DNA repair
• DNA transcription
• Chromatin structure

Research Areas

• Life sciences, Biophysics, Single molecule biophysics

Papers

• Solution AFM imaging and coarse-grained molecular modeling of yeast condensin structural variation coupled to the ATP hydrolysis cycle

  Hiroki Koide; Noriyuki Kodera; Shoji Takada; Tsuyoshi Terakawa

  bioRxiv, 03 Jan. 2025, Last author, Corresponding author

  
• Minimal protein list for chromatin replication-coupled symmetric histone recycling revealed throughin vitroreconstitution

  Fritz Nagae; Shizuko Endo; Yasuto Murayama; Tsuyoshi Terakawa

  bioRxiv, 19 Nov. 2024, Last author, Corresponding author

  
• SMC complex unidirectionally translocates DNA by coupling segment capture with an asymmetric kleisin path

  Masataka Yamauchi; Giovanni B. Brandani; Tsuyoshi Terakawa; Shoji Takada

  bioRxiv, 02 May 2024

  
• Molecular dynamics simulations of human cohesin subunits identify DNA binding sites and their potential roles in DNA loop extrusion

  Chenyang Gu; Shoji Takada; Giovanni B. Brandani; Tsuyoshi Terakawa

  PLOS Computational Biology, 04 Apr. 2025, Peer-reviewed, Corresponding author

  Access this paper :

  
• Molecular mechanism of parental H3/H4 recycling at a replication fork

  Fritz Nagae; Yasuto Murayama; Tsuyoshi Terakawa

  Nature Communications, 02 Nov. 2024, Peer-reviewed, Corresponding author

  
• Histone chaperone Nap1 dismantles an H2A/H2B dimer from a partially unwrapped nucleosome

  Fritz Nagae; Shoji Takada; Tsuyoshi Terakawa

  Nucleic Acids Research, 13 May 2023, Peer-reviewed, Corresponding author

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• ALS mutations in the TIA-1 prion-like domain trigger highly condensed pathogenic structures

  Naotaka Sekiyama; Kiyofumi Takaba; Saori Maki-Yonekura; Ken-ichi Akagi; Yasuko Ohtani; Kayo Imamura; Tsuyoshi Terakawa; Keitaro Yamashita; Daigo Inaoka; Koji Yonekura; Takashi S. Kodama; Hidehito Tochio

  Proceedings of the National Academy of Sciences, 16 Sep. 2022, Peer-reviewed

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• Coarse-grained molecular dynamics simulations of base-pair mismatch recognition protein MutS sliding along DNA

  Keisuke Inoue; Shoji Takada; Tsuyoshi Terakawa

  Biophysics and Physicobiology, 2022, Peer-reviewed, Corresponding author

  
• Single-molecule junction spontaneously restored by DNA zipper

  Takanori Harashima; Shintaro Fujii; Yuki Jono; Tsuyoshi Terakawa; Noriyuki Kurita; Satoshi Kaneko; Manabu Kiguchi; Tomoaki Nishino

  Nature Communications, 01 Oct. 2021, Peer-reviewed

  
• The lane-switch mechanism for nucleosome repositioning by DNA translocase

  Fritz Nagae; Giovanni B Brandani; Shoji Takada; Tsuyoshi Terakawa

  Nucleic Acids Research, 07 Aug. 2021, Peer-reviewed, Corresponding author

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• Modeling of DNA binding to the condensin hinge domain using molecular dynamics simulations guided by atomic force microscopy.

  Hiroki Koide; Noriyuki Kodera; Shveta Bisht; Shoji Takada; Tsuyoshi Terakawa

  PLoS computational biology, Jul. 2021, Peer-reviewed, Corresponding author

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• Defining the influence of Rad51 and Dmc1 lineage-specific amino acids on genetic recombination.

  Justin B Steinfeld; Ondrej Beláň; Youngho Kwon; Tsuyoshi Terakawa; Amr Al-Zain; Michael J Smith; J Brooks Crickard; Zhi Qi; Weixing Zhao; Rodney Rothstein; Lorraine S Symington; Patrick Sung; Simon J Boulton; Eric C Greene

  Genes & development, 01 Sep. 2019, Peer-reviewed

  
• Nucleosome Crowding in Chromatin Slows the Diffusion but Can Promote Target Search of Proteins.

  Ryo Kanada; Tsuyoshi Terakawa; Hiroo Kenzaki; Shoji Takada

  Biophysical Journal, 18 Jun. 2019, Peer-reviewed

  
• The condensin complex is a mechanochemical motor that translocates along DNA

  Tsuyoshi Terakawa; Shveta Bisht; Jorine M. Eeftens; Cees Dekker; Christian H. Haering; Eric C. Greene

  Science, Nov. 2017, Peer-reviewed, Lead author

  
• Sequential eviction of crowded nucleoprotein complexes by the exonuclease RecBCD molecular motor

  Tsuyoshi Terakawa; Sy Redding; Timothy D. Silverstein; Eric C. Greene

  Proceedings of the National Academy of Sciences, Aug. 2017, Peer-reviewed, Lead author

  
• How one-dimensional diffusion of transcription factors are affected by obstacles: coarse-grained molecular dynamics study

  Mami Saito; Tsuyoshi Terakawa; Shoji Takada

  Molecular Simulation, 2017, Peer-reviewed

  
• Dynamic Coupling among Protein Binding, Sliding, and DNA Bending Revealed by Molecular Dynamics

  Cheng Tan; Tsuyoshi Terakawa; Shoji Takada

  Journal of the American Chemical Society, Jul. 2016, Peer-reviewed

  
• Modeling Structural Dynamics of Biomolecular Complexes by Coarse-Grained Molecular Simulations

  Shoji Takada; Ryo Kanada; Cheng Tan; Tsuyoshi Terakawa; Wenfei Li; Hiroo Kenzaki

  Accounts of Chemical Research, Dec. 2015, Peer-reviewed

  
• p53 dynamics upon response element recognition explored by molecular simulations

  Tsuyoshi Terakawa; Shoji Takada

  Scientific Reports, Nov. 2015, Peer-reviewed, Lead author

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• Base triplet stepping by the Rad51/RecA family of recombinases

  Ja Yil Lee; Tsuyoshi Terakawa; Zhi Qi; Justin B. Steinfeld; Sy Redding; YoungHo Kwon; William A. Gaines; Weixing Zhao; Patrick Sung; Eric C. Greene

  Science, Aug. 2015, Peer-reviewed

  
• Multi-scale Ensemble Modeling of Modular Proteins with Intrinsically Disordered Linker Regions: Application to p53

  Tsuyoshi Terakawa; Junichi Higo; Shoji Takada

  Biophysical Journal, Aug. 2014, Peer-reviewed, Lead author

  
• RESPAC: Method to Determine Partial Charges in Coarse-Grained Protein Model and Its Application to DNA-Binding Proteins

  Tsuyoshi Terakawa; Shoji Takada

  Journal of Chemical Theory and Computation, Feb. 2014, Peer-reviewed, Lead author

  
• Energy landscape and multiroute folding of topologically complex proteins adenylate kinase and 2ouf-knot

  Wenfei Li; Tsuyoshi Terakawa; Wei Wang; Shoji Takada

  Proceedings of the National Academy of Sciences, Oct. 2012, Peer-reviewed

  
• p53 Searches on DNA by Rotation-Uncoupled Sliding at C-Terminal Tails and Restricted Hopping of Core Domains

  Tsuyoshi Terakawa; Hiroo Kenzaki; Shoji Takada

  Journal of the American Chemical Society, Sep. 2012, Peer-reviewed, Lead author

  
• Multiscale Ensemble Modeling of Intrinsically Disordered Proteins: p53 N-Terminal Domain

  Tsuyoshi Terakawa; Shoji Takada

  Biophysical Journal, Sep. 2011, Peer-reviewed, Lead author

  
• On Easy Implementation of a Variant of the Replica Exchange with Solute Tempering in GROMACS

  Tsuyoshi Terakawa; Tomoshi Kameda; Shoji Takada

  Journal of Computational Chemistry, May 2011, Peer-reviewed, Lead author

  

Misc.

• コンデンシンの分子モーター活性を同定

  寺川 剛

  生物物理, Apr. 2018, Peer-reviewed, Invited

  
• 染色体構造の形成をつかさどる分子モーター：コンデンシン

  寺川 剛

  実験医学, Feb. 2018, Invited

  
• 海外だより ～もし素人がアメリカで実験をはじめたら～

  寺川 剛

  生物物理, Mar. 2017, Invited

  
• コンデンシン複合体はDNAに沿って移動する メカノケミカルモーターである

  寺川 剛

  サイエンス誌に載った日本人研究者, 2017, Invited

  
• リコンビナーゼRad51/RecAファミリーによる 塩基トリプレット歩進

  寺川 剛

  サイエンス誌に載った日本人研究者, 2015, Invited

  

Presentations

• Molecular Dynamics of Histone Recycling in Chromatin Replication

  Tsuyoshi Terakawa

  The 47th Annual Meeting of the Molecular Biology Society of Japn, 27 Nov. 2024, Invited

  
• Reconstitution of chromatin replication-coupled symmetric histone recycling

  Fritz Nagae; Shizuko Endo; Yasuto Murayama; Tsuyoshi Terakawa

  5th East Asian Single-Molecule Biophysics Symposium, 08 Nov. 2024, Invited

  
• Biophysical approaches to the molecular mechanisms of histone recycling

  Tsuyoshi Terakawa

  第97回日本生化学会大会, 06 Nov. 2024, Invited

  
• Unveiling the molecular mechanism of histone recycling through biophysical approaches

  Tsuyoshi Terakawa

  21st IUPAB Congress 2024, 27 Jun. 2024, Invited

  
• Exploration of the molecular mechanism of histone recycling through molecular dynamics simulations, nanopore sequencing, and biochemical experiments

  Tsuyoshi Terakawa

  68th Annual Meeting of the Biophysical Society, Feb. 2024

  
• Biophysical approaches reveal molecular mechanism of histone recycling

  Tsuyoshi Terakawa

  The 46th Annual Meeting of the Molecular Biology Society of Japan, Dec. 2023, Invited

  
• Condensin’s journey on DNA with obstacles

  Tsuyoshi Terakawa

  4th East Asian Single-Molecule Biophysics Symposium, Sep. 2023, Invited

  
• トランスロケースとの衝突に伴うヌクレオソームの運命: エピゲノム継承の解明に向けて

  寺川 剛

  第15回エピジェネティクス研究会年会, May 2022, Invited

  
• Nucleosome repositioning dynamics upon collision with a translocase

  Tsuyoshi Terakawa

  The 44th Annual Meeting of the Molecular Biology Society of Japan, Dec. 2021, Invited

  
• DNA curtain assay of epigenome inheritance at single molecule level

  寺川 剛

  さきがけCREST「ゲノム合成」2020年度 合同領域会議, Jan. 2021, Invited

  
• Reconstitution of protein functions on chromatin curtain

  寺川 剛

  第43回分子生物学会年会ワークショップ「Reconstruction of the nucleus - Hierarchical understanding of the functional nucleus by reconstitutional approaches -」, Dec. 2020, Invited

  
• Toward the molecular mechanism of condensin translocation on DNA

  寺川 剛

  新学術領域「発動分子科学」第4回領域会議, Nov. 2020, Invited

  
• Histone modification and chromatin condensation explored by DNA curtain

  寺川 剛

  新学術領域「クロマチン潜在能」領域会議, May 2020, Invited

  
• DNA curtain assays and molecular dynamics simulations to elucidate the molecular mechanism of DNA transactions

  寺川 剛

  2nd ExCELLS Retreat for Young Scientists, Feb. 2020, Invited

  
• Condensinモーターの発動機構の解明に向けて

  寺川 剛

  新学術領域「発動分子科学」第2回領域会議, Jan. 2020

  
• Single-molecule observation of chromatin condensation induced by Condensin molecular motor

  寺川 剛

  第42回日本分子生物学会年会, Dec. 2019, Invited

  
• Single-molecule DNA curtain assays of epigenetic inheritance

  寺川 剛

  第２回キックオフCREST/さきがけ（ゲノム合成領域）合同領域会議, Nov. 2019, Invited

  
• DNAカーテン法を用いたDNA上の混み合い問題へのアプローチ

  寺川 剛

  OCU先端光科学シンポジウム, Oct. 2019, Invited

  
• DNA curtains: high-throughput single molecule imaging for DNA transactions

  寺川 剛

  第57回日本生物物理学会年会, Sep. 2019, Invited

  
• Condensin is a mechanochemical molecular motor that translocates along DNA

  寺川 剛

  理研エピジェネティクスセミナー, Sep. 2019, Invited

  
• A new type of DNA molecular motor: Condensin

  Tsuyoshi Terakawa

  2nd East Asian Single-Molecule Biophysics Symposium, Jul. 2019, Invited

  
• Relationship between histone modification and chromatin condensation explored by DNA curtain assay

  Tsuyoshi Terakawa

  新学術領域「クロマチン潜在能」 第2回領域会議, Jun. 2019, Invited

  
• 金・銀ナノ粒子を用いたコンデンシン分子モーターの超高分解能DNAカーテン測定

  寺川 剛

  新学術領域「発動分子科学」第2回領域会議, May 2019, Invited

  
• 染色体凝集の分子基盤: コンデンシン分子モーターの分子観察

  寺川 剛

  国立遺伝学研究所 研究集会「ゲノムの維持継承を支える分子基盤の包括的理解とその発展」, Feb. 2019, Invited

  
• DNA上の新しい分子モーター: コンデンシン

  寺川 剛

  分子モーター討論会, Nov. 2018, Invited

  
• The condensin complex is a mechanochemical molecular motor

  Tsuyoshi Terakawa

  The 56th Annual Meeting of The Biophysical Society of Japan, Sep. 2018, Invited

  
• The condensin complex is a new type of DNA molecular motor

  Tsuyoshi Terakawa

  The 79th Okazaki Conference, Sep. 2018, Invited

  
• The condensin complex is a mechanochemical motor that translocates along DNA

  Tsuyoshi Terakawa

  Genome Integrity Discussion Group (NYAS), Oct. 2017

  

External funds: Kakenhi

• クロマチンカーテン法によるクロマチン凝集の1分子蛍光顕微鏡観察

  Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)

  Biological Sciences

  Kyoto University

  寺川 剛

  From 01 Apr. 2021, To 31 Mar. 2023, Project Closed

  コンデンシン;分子モーター;染色体ループ形成機構;ヌクレオソーム;DNAカーテン法;一分子蛍光イメージング;蛍光顕微鏡観察;クロマチンカーテン;染色体凝集

  https://kaken.nii.ac.jp/grant/KAKENHI-PUBLICLY-21H00252/
• 金銀・ナノ粒子を用いたコンデンシン分子モーターの超高分解能DNAカーテン測定

  Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)

  Complex systems

  Kyoto University

  寺川 剛

  From 01 Apr. 2019, To 31 Mar. 2021, Project Closed

  コンデンシン;蛍光顕微鏡観察;DNAカーテン;一分子実験;量子ドット;金ナノ粒子;銀ナノ粒子;1分子蛍光顕微鏡観察;ステップサイズ測定

  https://kaken.nii.ac.jp/grant/KAKENHI-PUBLICLY-19H05392/
• DNAカーテン測定によるヒストン化学修飾がクロマチン凝集に与える影響の解明

  Grant-in-Aid for Scientific Research on Innovative Areas (Research in a proposed research area)

  Biological Sciences

  Kyoto University

  寺川 剛

  From 01 Apr. 2019, To 31 Mar. 2021, Project Closed

  コンデンシン;ヌクレオソーム;蛍光顕微鏡観察;DNAカーテン;一分子実験;染色体;ヒストン化学修飾;1分子蛍光顕微鏡観察;染色体凝集;クロマチン;エピジェネティクス

  https://kaken.nii.ac.jp/grant/KAKENHI-PUBLICLY-19H05260/
• Molecular mechanisms of a Condensin molecular motor explored by DNA curtain assays and coarse-grained molecular dynamics simulations

  Grant-in-Aid for Scientific Research (B)

  Basic Section 43040:Biophysics-related

  Kyoto University

  Tsuyoshi Terakawa

  From 01 Apr. 2019, To 31 Mar. 2022, Project Closed

  １分子蛍光イメージング;DNAカーテン;粗視化分子動力学シミュレーション;コンデンシン;蛍光顕微鏡観察;一分子実験;分子モーター

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-19H03194/
• Elucidation of the molecular mechanism of chromatin folding by single molecule experiments

  Grant-in-Aid for Research Activity Start-up

  0701:Biology at molecular to cellular levels, and related fields

  Kyoto University

  寺川 剛

  From 24 Aug. 2018, To 31 Mar. 2020, Project Closed

  DNAカーテン;蛍光顕微鏡;コンデンシン;染色体凝縮;分子モーター;染色体;ヌクレオソーム;全反射蛍光顕微鏡

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-18H06046/
• ナノポア解析と分子シミュレーションで探るヒストンリサイクリングの鎖バイアス機構

  Grant-in-Aid for Transformative Research Areas (B)

  Transformative Research Areas, Section (III)

  Kyoto University

  寺川 剛

  From 01 Apr. 2024, To 31 Mar. 2027, Granted

  ゲノム複製;エピゲノム継承;ヒストンリサイクリング;ナノポアシーケンシング;分子動力学シミュレーション

  https://kaken.nii.ac.jp/grant/KAKENHI-PLANNED-24H00883/
• エピゲノム継承への分子・細胞スケールブリッジアプローチの推進

  Grant-in-Aid for Transformative Research Areas (B)

  Transformative Research Areas, Section (III)

  Kyoto University

  寺川 剛

  From 01 Apr. 2024, To 31 Mar. 2027, Granted

  ヒストンリサイクリング;分子動力学シミュレーション;次世代シーケンシング;光ピンセット;染色体再構成

  https://kaken.nii.ac.jp/grant/KAKENHI-ORGANIZER-24H00882/
• Molecule- and cell-scale bridging approach to epigenome inheritance

  Grant-in-Aid for Transformative Research Areas (B)

  From 01 Apr. 2024, To 31 Mar. 2027

  https://kaken.nii.ac.jp/grant/KAKENHI-AREA-24B307/
• 細胞内複雑環境下の相分離による生体分子凝縮体形成の生物物理

  Grant-in-Aid for Scientific Research (B)

  Basic Section 43040:Biophysics-related

  Kyoto University

  高田 彰二

  From 01 Apr. 2024, To 31 Mar. 2027, Granted

  生体分子凝縮体;液液相分離;転写制御;シナプス可塑性

  https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-24K01991/

Education

Teaching subject(s)

• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IIC

  9027, Spring, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IIB

  9026, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IIA

  9025, Spring, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IID

  9028, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IA

  9021, Spring, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics ID

  9024, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IC

  9023, Spring, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminar on Theoretical Biophysics IB

  9022, Fall, Graduate School of Science, 1
• From 01 Apr. 2024, To 31 Mar. 2025

  Special Study CourseII(Biological Sciences) 23

  8726, Year-long, Faculty of Science, 12
• From 01 Apr. 2024, To 31 Mar. 2025

  Laboratory work in Biological Science A

  3762, Spring, Faculty of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Biomolecular Science I

  2716, Spring, Faculty of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Biomolecular Science II

  2715, Fall, Faculty of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Seminars in Biological Science B

  4732, Fall, Faculty of Science, 2
• From 01 Apr. 2024, To 31 Mar. 2025

  Advanced Course on Biological Science A

  9011, Spring, Graduate School of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IID

  9028, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IIC

  9027, Spring, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IIB

  9026, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IB

  9022, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IA

  9021, Spring, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IIA

  9025, Spring, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics ID

  9024, Fall, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminar on Theoretical Biophysics IC

  9023, Spring, Graduate School of Science, 1
• From 01 Apr. 2023, To 31 Mar. 2024

  Special Study CourseII(Biological Sciences) 23

  8726, Year-long, Faculty of Science, 12
• From 01 Apr. 2023, To 31 Mar. 2024

  Structural Biology

  8736, Fall, Faculty of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Seminars in Biological Science B

  4732, Fall, Faculty of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Laboratory work in Biological Science B

  3763, Spring, Faculty of Science, 2
• From 01 Apr. 2023, To 31 Mar. 2024

  Advanced Course on Biological Science B

  9012, Fall, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Practice of Basic Informatics (Faculty of Science)

  T003, Fall, Institute for Liberal Arts and Sciences, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IIB

  9026, Fall, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics ID

  9024, Fall, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IID

  9028, Fall, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IC

  9023, Spring, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Advanced Course on Biological Science B

  9012, Fall, Graduate School of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IB

  9022, Fall, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Special Study CourseII(Biological Sciences) 23

  8726, Year-long, Faculty of Science, 12
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IIC

  9027, Spring, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IIA

  9025, Spring, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Structural Biology

  8736, Fall, Faculty of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminar on Theoretical Biophysics IA

  9021, Spring, Graduate School of Science, 1
• From 01 Apr. 2022, To 31 Mar. 2023

  Laboratory work in Biological Science B

  3763, Spring, Faculty of Science, 2
• From 01 Apr. 2022, To 31 Mar. 2023

  Seminars in Biological Science B

  4732, Fall, Faculty of Science, 2
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IA

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IB

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IC

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics ID

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IIA

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IIB

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IIC

  Spring, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminar on Theoretical Biophysics IID

  Fall, 理学研究科
• From Apr. 2018, To Mar. 2019

  Seminars in Biological Science B

  Fall, 理学部
• From Apr. 2018, To Mar. 2019

  Laboratory work in Biological Science E

  Fall, 理学部
• From Apr. 2018, To Mar. 2019

  Special Study CourseII(Biological Sciences) 23

  Year-long, 理学部
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IA

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IB

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IC

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics ID

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IIA

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IIB

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IIC

  Spring, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminar on Theoretical Biophysics IID

  Fall, 理学研究科
• From Apr. 2019, To Mar. 2020

  Seminars in Biological Science B

  Fall, 理学部
• From Apr. 2019, To Mar. 2020

  Laboratory work in Biological Science E

  Fall, 理学部
• From Apr. 2019, To Mar. 2020

  Special Study CourseII(Biological Sciences) 23

  Year-long, 理学部
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IA

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IB

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IC

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics ID

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IIA

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IIB

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IIC

  Spring, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminar on Theoretical Biophysics IID

  Fall, 理学研究科
• From Apr. 2020, To Mar. 2021

  Seminars in Biological Science B

  Fall, 理学部
• From Apr. 2020, To Mar. 2021

  Laboratory Work in Biological Science E

  Fall, 理学部
• From Apr. 2020, To Mar. 2021

  Special Study CourseII(Biological Sciences) 23

  Year-long, 理学部
• From Apr. 2021, To Mar. 2022

  Structural Biology

  Fall, 理学部
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IA

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IB

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IC

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics ID

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IIA

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IIB

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IIC

  Spring, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminar on Theoretical Biophysics IID

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Seminars in Biological Science B

  Fall, 理学部
• From Apr. 2021, To Mar. 2022

  Laboratory Work in Biological Science E

  Fall, 理学部
• From Apr. 2021, To Mar. 2022

  Advanced Course on Biological Science B

  Fall, 理学研究科
• From Apr. 2021, To Mar. 2022

  Special Study CourseII(Biological Sciences) 23

  Year-long, 理学部

Administration

Faculty management (title, position)

• From 01 Apr. 2023, To 31 Mar. 2025

  理学研究科植物園運営委員会委員
• From 01 Apr. 2019, To 31 Mar. 2020

  理学研究科SACRA国際戦略部門委員会委員
• From 01 Apr. 2020, To 31 Mar. 2021

  理学研究科SACRA国際戦略部門委員会委員
• From 01 Apr. 2021, To 31 Mar. 2022

  理学研究科SACRA国際戦略部門委員会委員