Goto, Ryutaro(Field Science Education and Research Center (FSERC)) | Activity Database on Education and Research, Kyoto University

Goto, Ryutaro

Field Science Education and Research Center (FSERC) Assistant Professor

Last Updated :2025/04/29

Basic Information

Professional Memberships

From Aug. 2024
オーストラリア軟体動物学会

日本動物分類学会

日本貝類学会

日本進化学会

日本ベントス学会

日本動物学会

Academic Degree

Master of Human and Environmental Sciences(Kyoto University)

Ph.D, Human and Environmental Sciences(Kyoto University)

Research History

From Mar. 2017, To Present
Kyoto University, Field Science Education and Research Center, 助教

From May 2015, To Feb. 2017
ミシガン大学, 進化生物学・生態学部, 日本学術振興会海外特別研究員

From Apr. 2015, To May 2015
The University of Tokyo, Atmosphere and Ocean Research Institute, 海洋科学特定共同研究員

From Apr. 2012, To Mar. 2015
The University of Tokyo, Atmosphere and Ocean Research Institute, 日本学術振興会特別研究員（PD）

From Apr. 2009, To Mar. 2012
Kyoto University, 人間・環境学研究科, 日本学術振興会特別研究員（DC1）

Last Updated :2025/04/29

Research

Research Topics, Overview of the research

Research Topics
Marine Invertebrates, Ecology, Taxonomy, Evolutionary Biology, Molecular Phylogenetics, Symbiosis.
Overview of the research
My research focuses on evolution and diversification of marine invertebrates. I am especially interested in (1) how symbiotic species interactions affect morphological and ecological evolution of marine organisms and their community structure; and (2) how and why extreme sexual dimorphism evolved in various marine taxa.

Research Areas

Life sciences, Biodiversity and systematics

Life sciences, Ecology and environmental science

Papers

A symbiotic oyster in a shrimp burrow: phylogenetic position of Anomiostrea within the Ostreidae (Bivalvia)
Ryutaro Goto; Matteo Garzia; P Graham Oliver; Gustav Paulay; Daniele Salvi
Journal of Molluscan Studies, 30 Sep. 2024, Peer-reviewed, Lead author

紀伊半島南西部（和歌山県白浜町）殻のマゴコロガイの記録
後藤龍太郎; 柳下拓磨; 邉見由美
南紀生物, Jul. 2024, Lead author, Corresponding author

Within-host adaptive speciation of commensal yoyo clams leads to ecological exclusion, not co-existence
Teal A. Harrison; Ryutaro Goto; Jingchun Li; Diarmaid Ó Foighil
PeerJ, 05 Aug. 2024, Peer-reviewed
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Worms that repair: reconstruction of tubes is faster than normal growth in Spirobranchus akitsushima (Annelida: Serpulidae)
Genki Kobayashi; Ryutaro Goto
Marine Biology Research, Feb. 2024, Peer-reviewed, Last author
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Latitudinal shift of the associated hosts in Sagamiscintilla thalassemicola (Galeommatoidea: Galeommatidae), a rare ectosymbiotic bivalve that lives on the proboscis of echiuran worms
Ryutaro Goto; Taigi Sato; Hiroki Nakajima; Takahiro Sugiyama; Hiroshi Ishikawa
Journal of the Marine Biological Association of the United Kingdom, 06 Dec. 2023, Peer-reviewed, Lead author, Corresponding author

Living together in dead coral rocks: macrosymbiotic communities associated with Bonellia echiuran worms (Annelida: Thalassematidae: Bonelliinae), involving new commensal bivalve and amphipod species
Ryutaro Goto; Isao Hirabayashi; Koji Seike; Momo Yamashita; Michitaka Shimomura
Zoological Journal of the Linnean Society, 11 Oct. 2023, Peer-reviewed, Lead author, Corresponding author

Molecular and Morphological Assessment of Juvenile and Adult Forms in the Giant Worm Eunice Cf. Aphroditois (Annelida: Eunicidae) and Its Phylogenetic Position in the Family
Chiharu Shinomiya; Genki Kobayashi; Koji Seike; Momo Yamashita; Luna Yamamori; Takahiro Sugiyama; Mariko Kawamura; Kanto Nishikawa; Ryutaro Goto
Zoological Science, 24 Jul. 2023, Peer-reviewed, Last author

Records of the galeommatoid bivalve Nipponomysella subtruncata (Yokoyama, 1927), a commensal with the sipunculan Siphonosoma cumanense (Keferstein, 1867)from Miyagi and Aomori Prefectures, Japan
Hirokazu ABE; Genki KOBAYASHI; Takeru ODANAKA; Takashi OTA; Ryutaro GOTO; Takuya MINOKAWA; Masahiko WASHIO; Hirokazu ABE; Hiroaki FUKUMORI
石巻専修大学研究紀要, Mar. 2023

Evolutionary patterns of host switching, lifestyle mode, and the diversification history in symbiotic zoantharians
Hiroki Kise; Maria Eduarda Alves Santos; Chloé Julie Loïs Fourreau; Akira Iguchi; Ryutaro Goto; James Davis Reimer
Molecular Phylogenetics and Evolution, Feb. 2023, Peer-reviewed

Diversity, taxonomy, and evolution of symbiotic bivalves of the superfamily Galeommatoidea
Ryutaro Goto
The Chiribotan, Aug. 2022, Peer-reviewed, Invited, Lead author, Last author, Corresponding author
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Parallel chemical switches underlying pollinator isolation in Asian Mitella
T. Okamoto; Y. Okuyama; R. Goto; M. Tokoro; M. Kato
JOURNAL OF EVOLUTIONARY BIOLOGY, Mar. 2015, Peer-reviewed
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Patterns of shell utilization and preference in two sipunculan genera, Phascolion and Aspidosiphon
Kohei Oshiro; Akihiro Yoshikawa; Akira Asakura; Ryutaro Goto
Journal of the Marine Biological Association of the United Kingdom, 06 Jul. 2022, Peer-reviewed, Last author

Stasis and diversity in living fossils: Species delimitation and evolution of lingulid brachiopods
Ryutaro Goto; Tsuyoshi Takano; Koji Seike; Momo Yamashita; Gustav Paulay; Ku'ulei S. Rodgers; Cynthia L. Hunter; Piyoros Tongkerd; Shin'ichi Sato; Jae-Sang Hong; Kazuyoshi Endo
Molecular Phylogenetics and Evolution, Jul. 2022, Peer-reviewed, Lead author, Corresponding author

Nocturnal emission and post‐pollination change of floral scent in the leafflower tree, Glochidion rubrum, exclusively pollinated by seed‐parasitic leafflower moths
Tomoko Okamoto; Glenn P. Svensson; Ryutaro Goto; Atsushi Kawakita; Makoto Kato
Plant Species Biology, 18 Feb. 2022, Peer-reviewed

Giant spoon worms pumped out of their deep burrows: First collection of the main bodies of Ikeda taenioides (Annelida: Thalassematidae: Bonelliinae) in 88 years
Ryutaro Goto; Yumi Henmi; Yuto Shiozaki; Gyo Itani
Plankton and Benthos Research, 06 Aug. 2021, Peer-reviewed, Lead author, Corresponding author

Molecular phylogenetic assessment of Spirobranchus kraussii-complex (Annelida: Serpulidae) from the Japanese Archipelago
Genki Kobayashi; Ryutaro Goto
PeerJ, 14 Jul. 2021, Peer-reviewed, Last author
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Phylogeography of Mytilisepta virgata (Mytilidae: Bivalvia) in the northwestern Pacific: Cryptic mitochondrial lineages and mito-nuclear discordance
Yucheol Lee; Gang Ni; Jinkyung Shin; Taeho Kim; Elizabeth M.A. Kern; Yuseob Kim; Seung-Chul Kim; Benny Chan; Ryutaro Goto; Tomoyuki Nakano; Joong-Ki Park
Molecular Phylogenetics and Evolution, Apr. 2021, Peer-reviewed

Community-level plant–pollinator interactions in a palaeotropical montane evergreen oak forest ecosystem
Makoto Kato; Atsushi Kawakita; Ryutaro Goto; Tomoko Okamoto; Chisato Kobayashi; Yume Imada; Yuta Nakase; Tatsuki Nishioka; Bakham Chanthavong; Khamsing Keothumma; Yasuyuki Kosaka
Journal of Natural History, 26 Feb. 2021, Peer-reviewed

Snails riding mantis shrimps: Ectoparasites evolved from ancestors living as commensals on the host’s burrow wall
Ryutaro Goto; Tsuyoshi Takano; Douglas J. Eernisse; Makoto Kato; Yasunori Kano
Molecular Phylogenetics and Evolution, Feb. 2021, Peer-reviewed, Lead author, Corresponding author

Molecular and morphological systematics of the crinoid-parasitic snail genus Goodingia (Mollusca: Caenogastropoda: Eulimidae) with new insights into intrafamilial phylogenetic relationships
Tsuyoshi Takano; Ryutaro Goto
Marine Biodiversity, Feb. 2021, Peer-reviewed, Last author

Use of crustacean burrows as habitat by the marine snail Circulus cinguliferus (Gastropoda: Truncatelloidea: Vitrinellidae)
Ryutaro Goto; Taigi Sato
Plankton and Benthos Research, Feb. 2021, Peer-reviewed, Lead author, Corresponding author

Widening the host range of the ectosymbiotic scale-worm Asterophilia culcitae (Annelida: Polynoidae) to three echinoderm classes, with data on its body color variation
Takahiro Sugiyama; Naoto Jimi; Ryutaro Goto
Plankton and Benthos Research, 18 Nov. 2020, Peer-reviewed, Last author

Trophic segregation in a burrow: the stable carbon and nitrogen isotope ratios of the burrowing shrimp Upogebia major and its commensal bivalve Cryptomya busoensis.
Koji Seike; Ryutaro Goto
Plankton & Benthos Research, Sep. 2020, Peer-reviewed

Size and sex bias in air-exposure behavior during low tide of the intertidal hermit crab Clibanarius virescens (Krauss, 1843) (Decapoda: Anomura: Diogenidae)
Akihiro Yoshikawa; Ryutaro Goto; Chiaki I Yasuda; Akira Asakura
Journal of Crustacean Biology, 30 Mar. 2020, Peer-reviewed

Record of a rare mantis shrimp Bigelowina phalangium from the coasts of Shikoku facing the Seto Inland Sea, Japan
Shiozaki Yuto; Henmi Yumi; Goto Ryutaro; Itani Gyo; Ariyama Hiroyuki
CANCER, 2020, Peer-reviewed

Phylogeny of Echiura updated, with a revised taxonomy to reflect their placement in Annelida as sister group to Capitellidae
Ryutaro Goto; James Monnington; Marija Sciberras; Isao Hirabayashi; Greg W. Rouse
Invertebrate Systematics, 2020, Peer-reviewed, Lead author, Corresponding author

Records of the giant spoon worm Ikeda taenioides (Annelida: Echiura: Ikedidae) from the Kii Peninsula, the Sea of Japan, and Amami-Oshima Island
OSHIRO Kohei; HIRABAYASHI Isao; HENMI Yumi; GOTO Ryutaro
Japanese Journal of Benthology, 2020, Peer-reviewed, Last author

Worm-riding clam: description of Montacutona sigalionidcola sp. nov. (Bivalvia: Heterodonta: Galeommatidae) from Japan and its phylogenetic position
Ryutaro Goto; Makoto Tanaka
Zootaxa, Aug. 2019, Peer-reviewed

Remarkably loud snaps during mouth-fighting by a sponge-dwelling worm
Ryutaro Goto; Isao Hirabayashi; A. Richard Palmer
Current Biology, Jul. 2019, Peer-reviewed

Rediscovery of Ikedosoma elegans from Takasu tidal flat in Okayama
Luna Yamamori; Ryutaro Goto
Nanki Seibutsu, Jun. 2019

Morphology and Habitats of the Hermit-Crab-Associated Calyptraeid Gastropod Ergaea walshi
Yoshikawa Akihiro; Goto Ryutaro; Asakura Akira
Zoological science, Dec. 2018, Peer-reviewed

Evolutionary gain of red blood cells in a commensal bivalve (Galeommatoidea) as an adaptation to a hypoxic shrimp burrow
Ryutaro Goto; Hiroaki Fukumori; Yasunori Kano; Makoto Kato
Biological Journal of the Linnean Society, Sep. 2018, Peer-reviewed

An unusual habitat for bivalves: rediscovery of the enigmatic commensal clam Sagamiscintilla thalassemicola (Habe, 1962) (Bivalvia: Galeommatoidea) from spoon worm’s spoon
Ryutaro Goto; Hiroshi Ishikawa
Marine Biodiversity, 12 May 2018, Peer-reviewed

Molecular phylogeny of Maldanidae (Annelida): multiple losses of tube-capping plates and evolutionary shifts in habitat depth
Genki Kobayashi; Ryutaro Goto; Tsuyoshi Takano; Shigeaki Kojima
Molecular Phylogenetics and Evolution, May 2018, Peer-reviewed

Within-host speciation events in yoyo clams, obligate commensals with mantis shrimps, including one that involves a change in microhabitat and a loss of specialized traits
Ryutaro Goto; Teal A. Harrison; Diarmaid Ó Foighil
Biological Journal of the Linnean Society, May 2018, Peer-reviewed

First record of the rare spoon worm Ikedosoma elegans (Annelida: Echiura: Thalassematidae) from Shikoku Island, Japan
GOTO Ryutaro; HENMI Yumi; Corral Jonel Mangente; SHIOZAKI Yuto; KATO Testuya; ITANI Gyo
Japanese Journal of Benthology, 2018, Peer-reviewed

First record of Gymnogobius macrognathos in Akkeshi mud flat, Hokkaido, Japan and utilization of the Upogebia major burrow
Yumi Henmi; Ryutei Inui; Ryutaro Goto; Gyo Itani
Japanese Journal of Ichthyology, 2018, Peer-reviewed

Transfer of the gatekeeper sea anemone Verrillactis sp. (Cnidaria: Actiniaria: Sagartiidae) between shells by the host hermit crab Dardanus deformis (H. Milne Edwards, 1836) (Decapoda: Anomura: Diogenidae)
Yoshikawa Akihiro; Goto Ryutaro; Asakura Akira
Crustacean Research, 2018, Peer-reviewed

The enigmatic bivalve genus Paramya (Myoidea: Myidae): symbiotic association of an East Asian species with spoon worms (Echiura) and its transfer to the family Basterotiidae (Galeommatoidea)
Ryutaro Goto; Hiroshi Ishikawa; Yoichi Hamamura
JOURNAL OF THE MARINE BIOLOGICAL ASSOCIATION OF THE UNITED KINGDOM, Nov. 2017, Peer-reviewed

Combining in situ burrow casting and computed tomography scanning reveals burrow morphology and symbiotic associations in a burrow
Koji Seike; Ryutaro Goto
MARINE BIOLOGY, Mar. 2017, Peer-reviewed

Underwater observations of the giant spoon worm Ikeda taenioides (Annelida: Echiura: Ikedidae) in a subtidal soft-bottom environment in northeastern Japan, which survived tsunamis of the 2011 off the Pacific Coast of Tohoku Earthquake
Ryutaro Goto; Shingo Sakamoto; Jun Hayakawa; Koji Seike
JOURNAL OF OCEANOGRAPHY, Feb. 2017, Peer-reviewed

A new large tellinid species of the genus Pharaonella from the Ryukyu Archipelago, Japan (Mollusca, Bivalvia)
Makoto Kato; Luna Yamamori; Ryutaro Goto; Remi Tsubaki; Ken Ohsuga
ZOOKEYS, 2017, Peer-reviewed

Morphology, Biology, and Phylogenetic Position of the Bivalve Platomysia rugata (Heterodonta: Galeommatoidea), a Commensal with the Sipunculan Worm Sipunculus nudus
Ryutaro Goto; Hiroshi Ishikawa; Yoichi Hamamura
ZOOLOGICAL SCIENCE, Aug. 2016, Peer-reviewed

Symbiotic Association of the Bivalve Tellimya fujitaniana (Galeommatoidea) with the Heart Urchin Echinocardium cordatum (Spatangoida) in the Northwestern Pacific
Ryutaro Goto; Hiroshi Ishikawa; Yoichi Hamamura
ZOOLOGICAL SCIENCE, Aug. 2016, Peer-reviewed

A comprehensive molecular phylogeny of spoon worms (Echiura, Annelida): Implications for morphological evolution, the origin of dwarf males, and habitat shifts
Ryutaro Goto
MOLECULAR PHYLOGENETICS AND EVOLUTION, Jun. 2016, Peer-reviewed

Borniopsis mortoni sp n. (Heterodonta, Galeommatoidea, Galeommatidae sensu lato), a new bivalve commensal with a synaptid sea cucumber from Japan
Ryutaro Goto; Hiroshi Ishikawa
ZOOKEYS, 2016, Peer-reviewed

Coprinopsis asiaticiphlyctidospora sp nov., an agaric ammonia fungus from Amami and Okinawa, southern Japan
Toshimitsu Fukiharu; Kiminori Shimizu; Hideyuki Utsunomiya; Jay Kant Raut; Ryutaro Goto; Tomoko Okamoto; Makoto Kato; Reiko Horigome; Tatsuo Furuki; Noriko Kinjo
MYCOSCIENCE, Sep. 2014, Peer-reviewed

Mode of life of Anomiostrea coralliophila Habe, 1975 (Ostreidae): a symbiotic oyster living in ghost-shrimp burrows
Ryutaro Goto; Ken Ohsuga; Makoto Kato
JOURNAL OF MOLLUSCAN STUDIES, May 2014, Peer-reviewed

Evolution of symbiosis with Lingula (brachiopoda) in the bivalve superfamily Galeommatoidea (Heterodonta), with description of a new species of Koreamya
Ryutaro Goto; Hiroshi Ishikawa; Yoichi Hamamura; Shin'ichi Sato; Makoto Kato
JOURNAL OF MOLLUSCAN STUDIES, May 2014, Peer-reviewed

Active pollination favours sexual dimorphism in floral scent
Tomoko Okamoto; Atsushi Kawakita; Ryutaro Goto; Glenn P. Svensson; Makoto Kato
PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES, Dec. 2013, Peer-reviewed
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Molecular Phylogeny of Echiuran Worms (Phylum: Annelida) Reveals Evolutionary Pattern of Feeding Mode and Sexual Dimorphism
Ryutaro Goto; Tomoko Okamoto; Hiroshi Ishikawa; Yoichi Hamamura; Makoto Kato
PLOS ONE, Feb. 2013, Peer-reviewed

Molecular phylogeny of the bivalve superfamily Galeommatoidea (Heterodonta, Veneroida) reveals dynamic evolution of symbiotic lifestyle and interphylum host switching
Ryutaro Goto; Atsushi Kawakita; Hiroshi Ishikawa; Yoichi Hamamura; Makoto Kato
BMC EVOLUTIONARY BIOLOGY, Sep. 2012, Peer-reviewed

A novel brood-site pollination mutualism?: the root holoparasite Thonningia sanguinea (Balanophoraceae) and an inflorescence-feeding fly in the tropical rainforests of West Africa
Ryutaro Goto; Gen Yamakoshi; Tetsuro Matsuzawa
PLANT SPECIES BIOLOGY, May 2012, Peer-reviewed

いきものの不思議パートナーに制裁を加える植物 : 共生系が維持される仕組みを探る
川北篤; 後藤龍太郎
遺伝 : 生物の科学, Mar. 2012

Geographic mosaic of mutually exclusive dominance of obligate commensals in symbiotic communities associated with a burrowing echiuran worm
Ryutaro Goto; Makoto Kato
MARINE BIOLOGY, Feb. 2012, Peer-reviewed

A record of Satsuma (Luchuhadra) eucosmia (Pulmonata: Camaenidae) introduced in Amami-oshima Island
亀田勇一; 後藤龍太郎
ちりぼたん, Oct. 2011, Peer-reviewed

Morphological and Ecological Adaptation of Basterotia Bivalves (Galeommatoidea: Sportellidae) to Symbiotic Association with Burrowing Echiuran Worms
Ryutaro Goto; Yoichi Hamamura; Makoto Kato
ZOOLOGICAL SCIENCE, Mar. 2011, Peer-reviewed
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Mutualism favours higher host specificity than does antagonism in plant-herbivore interaction
Atsushi Kawakita; Tomoko Okamoto; Ryutaro Goto; Makoto Kato
PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES, Sep. 2010, Peer-reviewed

Selective flower abortion maintains moth cooperation in a newly discovered pollination mutualism
Ryutaro Goto; Tomoko Okamoto; E. Toby Kiers; Atsushi Kawakita; Makoto Kato
ECOLOGY LETTERS, Mar. 2010, Peer-reviewed

First Record of the Genus Megadenus Rosén, 1910 (Gastropoda: Eulimidae), Endoparasites of Sea Cucumbers, from Japan
Goto Ryutaro
Venus (Journal of the Malacological Society of Japan), 2010, Peer-reviewed

Chemical ecology of obligate pollination mutualisms: testing the 'private channel' hypothesis in the Breynia-Epicephala association
Glenn P. Svensson; Tomoko Okamoto; Atsushi Kawakita; Ryutaro Goto; Makoto Kato
NEW PHYTOLOGIST, 2010, Peer-reviewed

Obligate commensalism of Curvemysella paula (Bivalvia : Galeommatidae) with hermit crabs
Ryutaro Goto; Yoichi Hamamura; Makoto Kato
MARINE BIOLOGY, Jun. 2007, Peer-reviewed

Misc.

国際学会参加報告 Bivalve 2023–Where Are We Going＠ケンブリッジ大学に参加して
後藤龍太郎
日本ベントス学会誌, Jan. 2024, Lead author, Corresponding author

書評：新種発見物語
後藤龍太郎
日本進化学会ニュース, Jul. 2023, Lead author

海底の無脊椎動物の進化を紐解く
後藤龍太郎
日本進化学科ニュース, Mar. 2023, Invited, Lead author, Last author, Corresponding author

干潟の生物の多様性、共生、進化を探る
後藤龍太郎
教職員情報, Aug. 2021, Invited, Lead author

Acquisition of cryptic body coloration in a parasitic polychaete: does it steal pigments from its host?
杉山高大; 加山基; 宮下英明; 石川牧子; 朝倉彰; 後藤龍太郎
日本生態学会大会講演要旨(Web), 2021

Acquisition of cryptic body coloration in the parasitic scale worm Gastrolepidia clavigera Schmarda 1861
杉山高大; 加山基; 宮下英明; 石川牧子; 朝倉彰; 後藤龍太郎
日本ベントス学会・日本プランクトン学会合同大会講演要旨集, 2020

2019年日本ベントス学会・日本プランクトン学会合同大会自由集会2「環形動物の多様性と進化」開催報告
小林元樹; 杉山高大; 阿部博和; 中村真悠子; 自見直人; 後藤龍太郎
日本ベントス学会誌, 2020

Gastropod shells used by hermit crabs, as secondary habitats for various invertebrates
Yoshikawa Akihiro; Goto Ryutaro; Asakura Akira
CANCER, 2019, Invited

日本産ユムシ動物の多様性と系統分類(1)
後藤龍太郎
うみうし通信, Sep. 2012, Invited

Presentations

白鳳丸サンプルから明らかになった深海ユムシ類・ホシムシ類の多様性・進化
後藤龍太郎; 杉山高大; 四ノ宮千遥
北西太平洋の深海生物相白鳳丸KH-22-8、23-5次航海の成果を中心に, 19 Feb. 2025

海産無脊椎動物の多様性と進化
後藤龍太郎
日本学術会議「動物科学の最前線：めくるめく多様性を科学する（3）」, 15 Feb. 2025, Invited

An undescribed species of the genus Leucothoe Leach, 1814 (Amphipoda: Leucothoidae) associated with a bivalve collected from Sugashima, Mie Prefecture
花岡朋哉; 後藤龍太郎; 中島広喜; 小玉将史
日本甲殻類学会第62回大会

Living Inside Snapping Shrimp Burrows: Diversity and Phylogenetic Positions of Bivalves Commensal with Alpheid Shrimps in the Family Galeommatidae
Ryutaro Goto; Taigi Sato; Yumi Henmi; Tsuyoshi Takano; Takahiro Sugiyama; Gyo Itani
mollusc2024, Australia, 27 Nov. 2024

Diving into the shrimp hole: the hidden diversity and evolution of commensal galeommatid bivalves dwelling in snapping shrimp burrows
Ryutaro Goto; Taigi Sato; Yumi Henmi; Tsuyoshi Takano; Takahiro Sugiyama; Gyo Itani
The 5th Asian Marine Biology Symposium, Bangkok, Thailand, 2024, 29 Oct. 2024

Unveiling the species diversity of Phascolosoma (Annelida: Phascolosomatidae), sipunculan worms inhabiting hard substrates, in Japan
Chiharu Shinomiya; Luna Yamamori; Takahiro Sugiyama; Hiroshi Kajihara; Naoto Jimi; Genki Kobayashi; Akira Asakura; Michitaka Shimomura; Ryutaro Goto
The 5th Asian Marine Biology Symposium, Bangkok, Thailand, 28 Oct. 2024

スナギンチャク目(刺胞動物門:六放サンゴ綱)における宿主転換について
喜瀬浩輝; Maria Eduarda Alves Santos; Chloé Julie; Loïs Fourreau; 井口亮; 後藤龍太郎; James Davis Reimer
日本プランクトン学会・日本ベントス学会合同大会2024, 15 Sep. 2024

分子情報および外部形態に基づく日本産サメハダホシムシ類の種多様性の解明
四ノ宮千遥; 山守瑠奈; 杉山高大; 柁原宏; 自見直人; 小林元樹; 朝倉彰; 下村通誉; 後藤龍太郎
日本プランクトン学会・日本ベントス学会合同大会2024, 14 Sep. 2024

テッポウエビ類に巣穴共生するウロコガイ科二枚貝類の起源・進化・多様性
後藤龍太郎; 佐藤大義; 邉見由美; 髙野剛史; 杉山高大; 伊谷行
日本プランクトン学会・日本ベントス学会合同大会2024, 14 Sep. 2024

日本産サメハダホシムシ属（環形動物門：ホシムシ類サメハダホシムシ科）の種多様性
四ノ宮千遥; 山守瑠奈; 杉山高大; 柁原宏; 自見直人; 小林元樹; 朝倉彰; 下村通誉; 後藤龍太郎
日本動物分類学会第59回大会, Jun. 2024

Diversity and evolution of galeommatid clams inhabiting the burrows of snapping shrimps.
Ryutaro Goto; Taigi Sato; Yumi Henmi; Tsuyoshi Takano; Takahiro Sugiyama; Gyo Itani
令和６年度日本貝類学会大会, 14 Apr. 2024

Living together in the sea: Diversity and evolution of symbiotic bivalves and their host invertebrates
Ryutaro Goto
Bioblitz Guam 2024 symposium @ Marine Lab, University of Guam, 16 Feb. 2024, Invited

Down the shrimp hole: Evolution and diversity of galeommatid bivalves commensal with burrowing snapping shrimps
Ryutaro Goto; Taigi Sato; Yumi Henmi; Tsuyoshi Takano; Takahiro Sugiyama; Gyo Itani
BIVALVES – WHERE ARE WE GOING?, 08 Sep. 2023

Living together with spoon worms: Diversification and host specificity in commensal bivalves of Basterotiinae (Galeommatidae)
Ryutaro Goto; Taigi Sato; Isao Hirabayashi; Hiroshi Ishikawa
BIVALVES – WHERE ARE WE GOING?, 06 Sep. 2023

ユムシ共生性二枚貝の多様化様式と宿主特異性:ウロコガイ科イソカゼガイ亜科の分子系統解析
後藤龍太郎; 佐藤大義; 平林勲; 石川裕
2023年日本ベントス学会・日本プランクトン学会合同大会, 02 Sep. 2023

Host range of the parasitic polychaete Gastrolepidia clavigera
杉山高大; 朝倉彰; 後藤龍太郎
2023年日本ベントス学会・プランクトン学会合同大会, 01 Sep. 2023

超大型環形動物オニイソメ Eunice cf. aphroditois の体表から得られた日本初記録の小型寄生性カイアシ類
四ノ宮千遥; 後藤龍太郎; 幸塚久典; 山守瑠奈; 杉山高大; 上野大輔; 下村通誉
2023年日本プランクトン学会・ベントス学会合同大会, 01 Sep. 2023

海底生物の多様性、共生、音
後藤龍太郎
森里海連環学講座（パナソニック）, 19 Jul. 2023, Invited

Evolution of symbiosis between spoon worms (Annelida: Polychaeta: Thalassematidae) and their commensal bivalves
Ryutaro Goto; Isao Hirabayashi; Taigi Sato
14th International Polychaete Conference, 06 Jul. 2023

Which sea cucumbers do worms like? Host choice of the ectoparasitic polynoid Gastrolepidia clavigera Schmarda 1861
Takahiro Sugiyama; Akira Asakura; Ryutaro Goto
14th International Polychaete Conference, 04 Jul. 2023

Integrative taxonomy of Eunice cf. aphroditois (Annelida: Eunicidae) from Japan: comparative analysis of juvenile and adult forms and phylogenetic placement within the family
Chiharu Shinomiya; Genki Kobayashi; Koji Seike; Momo Yamashita; Luna Yamamori; Takahiro Sugiyama; Mariko Kawamura; Kanto Nishikawa; Ryutaro Goto
14th International Polychaete Conference, 04 Jul. 2023

超大型環形動物オニイソメ Eunice cf. aphroditois の体表に寄生する小型カイアシ類
四ノ宮千遥; 後藤龍太郎; 幸塚久典; 山守瑠奈; 杉山高大; 上野大輔; 下村通誉
日本動物分類学会第58回大会豊橋大会, 04 Jun. 2023

希少種ミドリユムシヤドリガイの琉球列島からの記録と新宿主
後藤龍太郎; 佐藤大義; 中島広喜; 杉山高大; 石川裕
日本動物分類学会第58回大会豊橋大会, 03 Jun. 2023

タテジマユムシ共生性ウロコムシ (環形動物門:多毛類) の分類と系統的位置
杉山高大; 自見直人; 朝倉彰; 後藤龍太郎
日本動物分類学会第58回大会豊橋大会, 03 Jun. 2023

海産底生生物の多様性と進化を探る
後藤龍太郎
2022年度第２回環境食品応用化学科セミナー（第６回光生命進化セミナー講演会）, 09 Dec. 2022, Invited

Exploring Evolutionary Trees of Marine Invertebrates
Ryutaro Goto
Virtual Guest Lecture (Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Surabaya), 07 Dec. 2022, Invited

ウロコガイ上科二枚貝類における矮雄の多様性と進化
後藤龍太郎; Diarmaid Ó Foighil
2022年日本プランクトン学会・日本ベントス学会合同大会, 03 Sep. 2022

環形動物門ウロコムシ科における生息基盤と類似した体色の進化
杉山高大; 朝倉彰; 後藤龍太郎
2022年日本プランクトン学会・日本ベントス学会合同大会, 03 Sep. 2022

海産大型多毛類オニイソメ (Eunice aphroditois)の幼体型および成体型の分子・形態情報に基づく比較検討
四ノ宮千遥; 小林元樹; 清家弘治; 山下桃; 西川完途; 後藤龍太郎
2022年日本プランクトン学会・日本ベントス学会合同大会, 03 Sep. 2022

Within-host ecological shift does not mean within-burrow coexistence for two bivalves commensal with mantis shrimp
Teal Harrison; Ryutaro Goto; Jingchun Li; Diarmaid Ó Foighil
World Congress of Malacology 2022, 05 Aug. 2022

海産無脊椎動物における寄生と共生：多様性、起源、進化を探る
後藤龍太郎
第2回ERATO共生進化機構先端セミナー, 09 Feb. 2022, Invited

沖縄産カンザシ種群から見つかった2系統および本州系統の遺伝的分化について
小林元樹; 後藤龍太郎
2021年日本ベントス学会・日本プランクトン学会合同大会, 19 Sep. 2021

寄生巻貝イシカワシタダミの起源
後藤龍太郎; 高野剛史; Douglas J. Eernisse; 加藤真; 狩野泰則
2021年日本ベントス学会・日本プランクトン学会合同大会, 19 Sep. 2021

オニイソメの分類と生態
四ノ宮千遥; 後藤龍太郎
2021年日本ベントス学会・日本プランクトン学会合同大会, 18 Sep. 2021

共生性ウロコムシ類のDNA バーコーディング分子系統解析
杉山高大; 自見直人; 小林元樹; 朝倉彰; 後藤龍太郎
2021年日本ベントス学会・日本プランクトン学会合同大会, 18 Sep. 2021

日本動物学会奨励賞受賞講演：海産無脊椎動物（特に軟体動物と環形動物）を対象とした進化生物学・系統分類学
後藤龍太郎
日本動物学会第92回オンライン米子大会, 03 Sep. 2021, Invited

干潟の地下で共に暮らす：共生貝類の多様性・生態・進化
後藤龍太郎
特別展普及講演会・日本貝類学会令和 3 年度大会公開講演会, 23 May 2021, Invited

Acquisition of cryptic body coloration in a parasitic polychaete: does it steal pigments from its host?
Takahiro SUGIYAMA; Motoki KAYAMA; Hideaki MIYASHITA; Makiko ISHIKAWA; Akira ASAKURA; Ryutaro GOTO
ESJ68, 18 Mar. 2021

Biodiversity in Sheltered Bays of Amami-Oshima Island, with a Focus on Benthoses and Symbioses in Tidal Flats
Ryutaro Goto
Sheltered Bay Environments of Amami-Oshima Island: Wonderful Biodiversity Hidden by Turbid Waters, 13 Mar. 2021, Invited

ウミシダに寄生するGoodingia属腹足類（ハナゴウナ科）の分類・分布・系統地理
高野剛史; 後藤龍太郎
令和二年度日本貝類学会, 05 Dec. 2020

Reconstruction of the flap-like structure at the opening of the dwelling tube by Spirobranchus kraussii (Annelida: Serpulidae)
Genki Kobayashi; Ryutaro Goto
2020年日本プランクトン・ベントス学会合同大会, 19 Sep. 2020

On the symbiont community associated with the burrows of Bonellia sp. (Annelida: Thalassematidae: Bonelliinae) inside the dead coral rocks
Ryutaro Goto; Isao Hirabayashi; Michitaka Shimomura
2020年日本プランクトン・ベントス学会合同大会, 19 Sep. 2020

Acquisition of cryptic body coloration in the parasitic scale worm Gasterolepidia clavigera Schmarda, 1861
Takahiro Sugiyama; Motoki Kayama; Hideaki Miyashita; Makiko Ishikawa; Ryutaro Goto
日本プランクトン・ベントス学会合同大会2020, 19 Sep. 2020

ゴカイも音を出す：攻撃行動に伴う発音とそのメカニズム
後藤龍太郎
名古屋大学アドバンス生命理学特論：GTR Seminar on Zoom, 16 Jul. 2020, Invited

キムラハナカゴオトヒメゴカイ（環形動物門：オトヒメゴカイ科）の種内闘争の際の発音とそのメカニズム
後藤龍太郎
三崎談話会第299回, 04 Dec. 2019, Invited

Loud snaps by a tiny marine worm and its sound production mechanism
Ryutaro Goto; Isao Hirabayashi; A. Richard Palmer
The 6th Annual Meeting of the Society for Bioacoustics, 22 Nov. 2019, Invited

Diversity of bivalves symbiotic with crustaceans
Ryutaro Goto
The Fourth Asian Marine Biology Symposium, 05 Nov. 2019, Invited

Morphological and molecular assessment of the species diversity of the genus Phascolion (Annelida: Sipuncula) in Japan
Kohei Oshiro; Akihiro Yoshikawa; Genki Kobayashi; Ryutaro Goto
The Foruth Asian Marine Biology Symposium, 05 Nov. 2019

Interspecific or intraspecific variation?: host-specific color morphs in the parasitic scale worm Gastrolepida clavigera (Annelida: Polynoidae)
Takahiro Sugiyama; Ryutaro Goto
The Fourth Asian Marine Biology Symposium, 05 Nov. 2019

Size biased landing behavior of the intertidal hermit crab Clibanarius virescens
Akihiro Yoshikawa; Ryutaro Goto; Chiaki Yasuda; Akira Asakura
Annual Meeting of Japanese Association of Benthology, 20 Sep. 2019

Species diversity of shell-dwelling sipunculans
Kohei Oshiro; Akihiro Yoshikawa; Genki Kobayashi; Ryutaro Goto
Annual Meeting of Japanese Association of Benthology, 19 Sep. 2019

Discovery of Asterophilia from holothurians
Takahiro Sugiyama; Ryutaro Goto
Annual Meeting of Japanese Association of Benthology, 19 Sep. 2019

Sound production associated with a mouth attack in Leocratides kimuraorum
Ryutaro Goto; Isao Hirabayashi; A. Richard Palmer
Annual Meeting of Japanese Association of Benthology, 18 Sep. 2019

Host-specific color morphs in the scale worm Gastrolepidia clavigera
Takahiro Sugiyama; Ryutaro Goto
Annual Meeting of Japanese Association of Benthology, 18 Sep. 2019

Morphological and ecological evolutionary pattern of Maldanidae
Genki Kobayashi; Ryutaro Goto; Tsuyoshi Takano; Shigeaki Kojima
Annual Meeting of Japanese Association of Benthology, 18 Sep. 2019

Evolution of yoyo clams: within-host ecological shifts lead to dramatic morphological changes
Ryutaro Goto; Teal A. Harrison; Diarmaid Ó Foighil
World Congress of Malacology 2019, 15 Apr. 2019

Morphological adaptation and habitat preference in the slipper limpet Ergaea walshi (Gastropoda: Calyptraeidae), a facultative associate with hermit crab living inside gastropod shells used by hermit crabs.
Yoshikawa Akihiro; Ryutaro Goto; Akira Asakura
Symposium of Integrative Biology: Biodiversity in Asia, 21 Feb. 2019

Diversity and evolution of spoon worms and their associated fauna
Ryutaro Goto
Symposium of Integrative Biology: Biodiversity in Asia, 21 Feb. 2019, Invited

Evolutionary shifts in habitat depths of bamboo worms (Annelida: Maldanidae)
Genki Kobayashi; Ryutaro Goto; Tsuyoshi Takano; Shigeaki Kojima
15th Deep-Sea Biology Symposium, 10 Sep. 2018

干潟で見られる共生関係の多様性と進化
後藤龍太郎
2018年度日本プランクトン学会・日本ベントス学会・合同大会, 10 Sep. 2018, Invited

ヒラフネガイのヤドカリ貝殻内部への共生様式
吉川晟弘; 後藤龍太郎; 朝倉彰
2018年度日本プラントン学会・日本ベントス学会・合同大会, 10 Sep. 2018

貝殻を背負って生きるホシムシ類の貝殻利用様式
大城匡平; 吉川晟弘; 後藤龍太郎
2018年度日本プランクトン学会・ベントス学会・合同大会, 10 Sep. 2018

Transfer of the sea anemone Verrillactis sp. by a host hermit crab
Akihiro Yoshikawa; Ryutaro Goto; Akira Asakura
NCB, 20 May 2018

How do seven commensal bivalves share the same host?
Teal Harrison; Ryutaro Goto; Michael J Boyle; Diarmaid Ó Foighil
SICB annual meeting, 04 Jan. 2018

ヤドカリに片利共生するヒラフネガイの宿主利用形式と形態適応
吉川晟弘; 後藤龍太郎; 朝倉彰
日本動物学会, Sep. 2017

潮間帯ヤドカリに見られる日光浴行動の効用
吉川晟弘; 後藤龍太郎; 朝倉彰
日本ベントス学会, Sep. 2017

ウロコガイ科二枚貝における住み込み共生と進化
後藤龍太郎
日本ベントス学会, Sep. 2017, Invited

Living together with crustaceans: Evolution and adaption of symbiotic mollusks
Ryutaro Goto
International Symposium on evolutionary biology of parasitic and symbiotic relationships between molluscs and crustaceans, Apr. 2017, Invited

住み込み共生する二枚貝の進化と適応
後藤龍太郎
生態研セミナー, 2017, Invited

腕足動物シャミセンガイ類の多様性と分子系統解析
後藤龍太郎; Eric Simon; 遠藤一佳
日本動物分類学会第53回大会, 2017

日本産タケフシゴカイ科環形動物の分子系統
小林元樹; 後藤龍太郎; 高野剛史; 小島茂明
日本動物分類学会第53回大会, 2017

Epifaunal life style and surface deposit feeding may enable bamboo worms (Annelida: Maldanidae) to invade chemosynthesis-based ecosystems
Kobayashi Genki; Ryutaro Goto; Tsuyoshi Takano; Shigeaki Kojima
6th International Symposium on Chemosynthesis-Based Ecosystems, 2017

Possibility and Challenges of Emergent Workshop: Co-designing the ecosystems of 2030 with future generations
Minori TOKITO; Daisuke AKAISHI; Juri HORI; Naoko TOKUCHI; Ryunosuke TATENO; Ryutaro GOTO; Keita SUZUKI; Nao SAKANOUE; Kodzue KINOSHITA; Akira ASAKURA
ESJ69, 15 Mar. 2022

Books and Other Publications

保全上重要なわかやまの自然－和歌山県レッドデータブック－2022年改訂版
後藤龍太郎, Contributor, スナウロコムシヤドリガイ;ミドリシャミセンガイ類似種;サナダユムシ
Nov. 2022

海産無脊椎動物多様性学: 100年の歴史とフロンティア
京都大学フィールド科学教育研究センター瀬戸臨海実験所創立100周年記念出版編集委員会, 第3部総論海洋環境における種内・種間の相互作用; 3.3 巣穴共生住み込み共生が創出する生物多様性─ユムシ類の共生生物群集
京都大学学術出版会, 21 Nov. 2022
Find Kyoto University's collection of books :

海岸動物の生態学入門ベントスの多様性に学ぶ
日本ベントス学会, Contributor, 共生二枚貝の宿主転換と多様化
海文堂出版, Sep. 2020

Obligate Pollination Mutualism. Ecological Research Monograph
Ryutaro Goto, Contributor, Role of Selective Flower Abortion in the Maintenance of Obligate Pollination Mutualism
Springer, Tokyo, 2017, Not refereed

Species Diversity of Animals in Japan
Ryutaro Goto, Contributor, The Echiura of Japan: Diversity, Classification, Phylogeny, and Their Associated Fauna
Springer, 2016, Not refereed

奄美群島の自然史学南西島嶼の生物多様性
後藤龍太郎, Contributor, 第６章｜居候して暮らす–南西諸島の干潟における共生二枚貝類の多様性
東海大学出版会, 2016, Not refereed

Awards

17 Apr. 2024
科学技術分野の文部科学大臣表彰若手科学者賞(令和6年度)

17 Apr. 2024
令和６年度科学技術分野の文部科学大臣表彰若手科学者賞

05 Aug. 2022
第２２回日本進化学会研究奨励賞

21 Apr. 2013
The Malacological Society of Japan, Young Scientist Award

10 Sep. 2018
日本ベントス学会, 日本ベントス学会奨励賞

03 Sep. 2021
日本動物学会, 日本動物学会奨励賞

24 Oct. 2021
Carcinological Society of Japan, Excellent Paper Award

Media Coverage

会誌「南紀生物」発行南紀生物同好会
紀伊民報, 29 Jun. 2024, Paper

二枚貝、ヨコエビ、新種発見
読売新聞, 17 Nov. 2023, Paper

絶滅したものや絶滅の恐れのある和歌山の海に住む生き物たちの展示京都大学白浜水族館
テレビ和歌山, 6wakaイブニング, 12 Mar. 2024, Media report

絶滅種や危惧種、84点の標本紹介京大白浜水族館で特別展 4月7日まで／和歌山
毎日新聞, 28 Jan. 2024, Paper

絶滅の恐れある海の生物／標本やパネルで紹介／京大白浜水族館
紀伊民報, 26 Jan. 2024, Paper

希少な二枚貝を採取／京大など／串本の海で４２年ぶり
紀伊民報, 24 Jan. 2024, Paper

新種の貝とヨコエビ和歌山・串本沖で京大が発見、水族館で展示
毎日新聞, 27 Oct. 2023, Paper

External funds: Kakenhi

Origins and evolution of dwarf males in marine bivalves
Grant-in-Aid for Early-Career Scientists
Basic Section 45030:Biodiversity and systematics-related
Kyoto University
Ryutaro Goto
From 01 Apr. 2020, To 31 Mar. 2023, Project Closed
性的二形;矮雄;進化;共生;繁殖集団サイズ;分子系統解析;二枚貝;性的二型;軟体動物門
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-20K15860/

Mechanism underling species radiation associated with the loss of symbiotic habit in marine clams
Grant-in-Aid for Research Activity Start-up
Kyoto University
Ryutaro Goto
From 25 Aug. 2017, To 31 Mar. 2019, Project Closed
二枚貝;種分化;共生;多様化;ウロコガイ科;進化;系統解析;ウロコガイ上科
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-17H06795/

Biogeography and evolution of gastropods in deep-sea chemosynthetic environments
Grant-in-Aid for Scientific Research (B)
The University of Tokyo
Yasunori Kano
From 01 Apr. 2015, To 31 Mar. 2019, Project Closed
化学合成群集;分子系統樹;生物地理;深海;幼生;比較解剖;化石;古生態;系統進化;海洋生態;層位・古生物学;解剖学;分類学
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-15H04412/

Diversification driven by within-host speciation in marine symbiotic mollusks
Grant-in-Aid for Scientific Research (C)
Basic Section 45030:Biodiversity and systematics-related
Kyoto University
後藤龍太郎
From 01 Apr. 2023, To 31 Mar. 2026, Granted
種分化;多様化;共生;軟体動物;ニッチ分化
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23K05906/

植物と送粉者の種特異性における化合物のブレンドとキラリティーの役割
Grant-in-Aid for Scientific Research (B)
Basic Section 45040:Ecology and environment-related
Gifu University
岡本朋子
From 01 Apr. 2021, To 31 Mar. 2025, Granted
キラリティ;種特異性;花の匂い;送粉共生;誘引;送粉;種分化;送粉共生系;ブレンド
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-21H02562/

植物と送粉者の種特異性における化合物のブレンドとキラリティーの役割
Grant-in-Aid for Scientific Research (B)
Basic Section 45040:Ecology and environment-related
Gifu University
岡本朋子
From 01 Apr. 2021, To 31 Mar. 2025, Granted
キラリティ;種特異性;花の匂い;送粉共生;誘引;送粉;種分化;送粉共生系;ブレンド
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-23K21339/

External funds: others

公益財団法人京都大学教育研究振興財団, 国際研究集会発表助成
From 01 Jul. 2023, To 09 Jul. 2023
後藤龍太郎

平成29年度第I期京都大学若手研究者スタートアップ研究費
From Jun. 2017, To Aug. 2017
後藤龍太郎

いしずえ
From 01 Apr. 2019, To 31 Mar. 2020
後藤龍太郎

公益財団法人京都大学教育研究振興財団研究活動推進助成
From 01 Apr. 2019, To 31 Mar. 2020
後藤龍太郎

Last Updated :2025/04/29

Education

Teaching subject(s)

From 01 Apr. 2024, To 31 Mar. 2025
ILAS Seminar
Z001, Spring, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2024, To 31 Mar. 2025
Field Study on Connectivity of Hills, Humans and Oceans IV
Y306, Fall, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2024, To 31 Mar. 2025
Marine biology with laboratory & field work 1
4734, Spring, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Seminars in Biological Science B
4732, Fall, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Invertebrate Zoology
2708, Fall, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Biological Oceanography
8732, Spring, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Marine biology with laboratory & field work 4
4737, Spring, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Marine biology with laboratory & field work 2
4735, Spring, Faculty of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Advanced Lectures : Systematic Marine Biology
8040, Spring, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Advanced Lectures : Integrative Biological Diversity
9901, Spring, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Seminar on Marine Biology A
8133, Spring, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Advanced Lectures: Functional Morphology and Phylogenetics B
8132, Fall, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Advanced Lectures: Functional Morphology and Phylogenetics A
8131, Spring, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Seminar on Marine Biology D
8136, Fall, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Seminar on Marine Biology C
8135, Spring, Graduate School of Science, 2

From 01 Apr. 2024, To 31 Mar. 2025
Seminar on Marine Biology B
8134, Fall, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
ILAS Seminar
Z001, Spring, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2023, To 31 Mar. 2024
Field Study on Connectivity of Hills, Humans and Oceans IV
Y306, Fall, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2023, To 31 Mar. 2024
Advanced Lectures : Phylogenetic analysis of invertebrates B
8130, Fall, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Advanced Lectures : Phylogenetic analysis of invertebrates A
8129, Spring, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Advanced Lectures : Systematic Marine Biology
8040, Spring, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Seminar on Marine Biology A
8133, Spring, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Seminar on Marine Biology D
8136, Fall, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Seminar on Marine Biology C
8135, Spring, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Seminar on Marine Biology B
8134, Fall, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Advanced Lectures : Integrative Biological Diversity
9901, Spring, Graduate School of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Marine biology with laboratory & field work 1
4734, Spring, Faculty of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Seminars in Biological Science B
4732, Fall, Faculty of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Invertebrate Zoology
2708, Fall, Faculty of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Biological Oceanography
8732, Spring, Faculty of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Marine biology with laboratory & field work 4
4737, Spring, Faculty of Science, 2

From 01 Apr. 2023, To 31 Mar. 2024
Marine biology with laboratory & field work 2
4735, Spring, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Seminar on Marine Biology C
8135, Spring, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Advanced Lectures: Functional Morphology and Phylogenetics B
8132, Fall, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Marine biology with laboratory & field work 4
4737, Spring, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Marine biology with laboratory & field work 2
4735, Spring, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Marine biology with laboratory & field work 1
4734, Spring, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Field Study on Connectivity of Hills, Humans and Oceans IV
Y306, Fall, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2022, To 31 Mar. 2023
Seminars in Biological Science B
4732, Fall, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Seminar on Marine Biology D
8136, Fall, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Seminar on Marine Biology A
8133, Spring, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
ILAS Seminar
Z001, Spring, Institute for Liberal Arts and Sciences, 2

From 01 Apr. 2022, To 31 Mar. 2023
Seminar on Marine Biology B
8134, Fall, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Invertebrate Zoology
2708, Fall, Faculty of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Advanced Lectures: Functional Morphology and Phylogenetics A
8131, Spring, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Advanced Lectures : Systematic Marine Biology
8040, Spring, Graduate School of Science, 2

From 01 Apr. 2022, To 31 Mar. 2023
Biological Oceanography
8732, Spring, Faculty of Science, 2

From Apr. 2017, To Mar. 2018
Advanced Lectures : Phylogenetic analysis of invertebrates A
Spring, 理学研究科

From Apr. 2017, To Mar. 2018
Advanced Lectures : Phylogenetic analysis of invertebrates B
Fall, 理学研究科

From Apr. 2017, To Mar. 2018
Biological Oceanography
Spring, 理学部

From Apr. 2017, To Mar. 2018
Seminar on Marine Biology A
Spring, 理学研究科

From Apr. 2017, To Mar. 2018
Seminar on Marine Biology B
Fall, 理学研究科

From Apr. 2017, To Mar. 2018
Seminar on Marine Biology C
Spring, 理学研究科

From Apr. 2017, To Mar. 2018
Seminar on Marine Biology D
Fall, 理学研究科

From Apr. 2017, To Mar. 2018
Advanced Lectures : Systematic Marine Biology
Spring, 理学研究科

From Apr. 2017, To Mar. 2018
Invertebrate Zoology
Fall, 理学部

From Apr. 2017, To Mar. 2018
Marine biology with laboratory & field work 1
Spring, 理学部

From Apr. 2017, To Mar. 2018
Marine biology with laboratory & field work 2
Spring, 理学部

From Apr. 2017, To Mar. 2018
Marine biology with laboratory & field work 4
Spring, 理学部

From Apr. 2018, To Mar. 2019
ILAS Seminar
Spring, 全学共通科目

From Apr. 2018, To Mar. 2019
Field Study on Connectivity of Hills, Humans and Oceans IV
Fall, 全学共通科目

From Apr. 2018, To Mar. 2019
Advanced Lectures: Functional Morphology and Phylogenetics A
Spring, 理学研究科

From Apr. 2018, To Mar. 2019
Advanced Lectures: Functional Morphology and Phylogenetics B
Fall, 理学研究科

From Apr. 2018, To Mar. 2019
Biological Oceanography
Spring, 理学部

From Apr. 2018, To Mar. 2019
Seminar on Marine Biology A
Spring, 理学研究科

From Apr. 2018, To Mar. 2019
Seminar on Marine Biology B
Fall, 理学研究科

From Apr. 2018, To Mar. 2019
Seminar on Marine Biology C
Spring, 理学研究科

From Apr. 2018, To Mar. 2019
Seminar on Marine Biology D
Fall, 理学研究科

From Apr. 2018, To Mar. 2019
Advanced Lectures : Systematic Marine Biology
Spring, 理学研究科

From Apr. 2018, To Mar. 2019
Invertebrate Zoology
Fall, 理学部

From Apr. 2018, To Mar. 2019
Seminars in Biological Science B
Fall, 理学部

From Apr. 2018, To Mar. 2019
Inter Lab Studies, Biological Science
Spring, 理学研究科

From Apr. 2018, To Mar. 2019
Marine biology with laboratory & field work 1
Spring, 理学部

From Apr. 2018, To Mar. 2019
Marine biology with laboratory & field work 2
Spring, 理学部

From Apr. 2018, To Mar. 2019
Marine biology with laboratory & field work 4
Spring, 理学部

From Apr. 2017, To Mar. 2018
Field Study on Connectivity of Hills, Humans and OceansIV
Fall, 全学共通科目

From Apr. 2017, To Mar. 2018
Inter Lab Studies, Biological Science
Spring, 理学研究科

From Apr. 2019, To Mar. 2020
ILAS Seminar
Spring, 全学共通科目

From Apr. 2019, To Mar. 2020
Advanced Lectures : Phylogenetic analysis of invertebrates A
Spring, 理学研究科

From Apr. 2019, To Mar. 2020
Advanced Lectures : Phylogenetic analysis of invertebrates B
Fall, 理学研究科

From Apr. 2019, To Mar. 2020
Field Study on Connectivity of Hills, Humans and Oceans IV
Fall, 全学共通科目

From Apr. 2019, To Mar. 2020
Biological Oceanography
Spring, 理学部

From Apr. 2019, To Mar. 2020
Seminar on Marine Biology A
Spring, 理学研究科

From Apr. 2019, To Mar. 2020
Seminar on Marine Biology B
Fall, 理学研究科

From Apr. 2019, To Mar. 2020
Seminar on Marine Biology C
Spring, 理学研究科

From Apr. 2019, To Mar. 2020
Seminar on Marine Biology D
Fall, 理学研究科

From Apr. 2019, To Mar. 2020
Advanced Lectures : Systematic Marine Biology
Spring, 理学研究科

From Apr. 2019, To Mar. 2020
Invertebrate Zoology
Fall, 理学部

From Apr. 2019, To Mar. 2020
Seminars in Biological Science B
Fall, 理学部

From Apr. 2019, To Mar. 2020
Marine biology with laboratory & field work 1
Spring, 理学部

From Apr. 2019, To Mar. 2020
Marine biology with laboratory & field work 2
Spring, 理学部

From Apr. 2019, To Mar. 2020
Marine biology with laboratory & field work 4
Spring, 理学部

From Apr. 2020, To Mar. 2021
ILAS Seminar :The Sea is longing for the forest
Spring, 全学共通科目

From Apr. 2020, To Mar. 2021
Advanced Lectures: Functional Morphology and Phylogenetics A
Spring, 理学研究科

From Apr. 2020, To Mar. 2021
Advanced Lectures: Functional Morphology and Phylogenetics B
Fall, 理学研究科

From Apr. 2020, To Mar. 2021
Field Study on Connectivity of Hills, Humans and Oceans IV : Effects on coastal ecosystem from land, river and human ...
Fall, 全学共通科目

From Apr. 2020, To Mar. 2021
Biological Oceanography
Spring, 理学部

From Apr. 2020, To Mar. 2021
Seminar on Marine Biology A
Spring, 理学研究科

From Apr. 2020, To Mar. 2021
Seminar on Marine Biology B
Fall, 理学研究科

From Apr. 2020, To Mar. 2021
Seminar on Marine Biology C
Spring, 理学研究科

From Apr. 2020, To Mar. 2021
Seminar on Marine Biology D
Fall, 理学研究科

From Apr. 2020, To Mar. 2021
Advanced Lectures : Systematic Marine Biology
Spring, 理学研究科

From Apr. 2020, To Mar. 2021
Invertebrate Zoology
Fall, 理学部

From Apr. 2020, To Mar. 2021
Seminars in Biological Science B
Fall, 理学部

From Apr. 2020, To Mar. 2021
Marine Biology with Laboratory & Field Work 1
Spring, 理学部

From Apr. 2020, To Mar. 2021
Marine Biology with Laboratory & Field Work 2
Spring, 理学部

From Apr. 2020, To Mar. 2021
Marine Biology with Laboratory & Field Work 4
Spring, 理学部

From Apr. 2021, To Mar. 2022
ILAS Seminar :The Sea is longing for the forest
Spring, 全学共通科目

From Apr. 2021, To Mar. 2022
Advanced Lectures : Phylogenetic analysis of invertebrates A
Spring, 理学研究科

From Apr. 2021, To Mar. 2022
Advanced Lectures : Phylogenetic analysis of invertebrates B
Fall, 理学研究科

From Apr. 2021, To Mar. 2022
Field Study on Connectivity of Hills, Humans and Oceans IV : Effects on coastal ecosystem from land, river and human ...
Fall, 全学共通科目

From Apr. 2021, To Mar. 2022
Biological Oceanography
Spring, 理学部

From Apr. 2021, To Mar. 2022
Seminar on Marine Biology A
Spring, 理学研究科

From Apr. 2021, To Mar. 2022
Seminar on Marine Biology B
Fall, 理学研究科

From Apr. 2021, To Mar. 2022
Seminar on Marine Biology C
Spring, 理学研究科

From Apr. 2021, To Mar. 2022
Seminar on Marine Biology D
Fall, 理学研究科

From Apr. 2021, To Mar. 2022
Advanced Lectures : Systematic Marine Biology
Spring, 理学研究科

From Apr. 2021, To Mar. 2022
Invertebrate Zoology
Fall, 理学部

From Apr. 2021, To Mar. 2022
Seminars in Biological Science B
Fall, 理学部

From Apr. 2021, To Mar. 2022
Marine Biology with Laboratory & Field Work 1
Spring, 理学部

From Apr. 2021, To Mar. 2022
Marine Biology with Laboratory & Field Work 2
Spring, 理学部

From Apr. 2021, To Mar. 2022
Marine Biology with Laboratory & Field Work 4
Spring, 理学部

Student achievements: Awards

日本ベントス学会2022年度大会学生発表賞
Chiharu Shinomiya, 日本ベントス学会, 21 Sep. 2022

Last Updated :2025/04/29

Administration

Faculty management (title, position)

From 01 Apr. 2017
研究プログラム委員会　委員

From 01 Apr. 2017
社会連携委員会　委員

From 01 Apr. 2017
図書委員会　委員

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Goto, Ryutaro

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